Carcharodus

Carcharodus Hübner, [1819]

Carcharodus Hübner, [1819]; Verz. bek. Schmett. (7): 110; TS: Papilio alceae Esper
Reverdinus Ragusa, 1919; Natural. Sicil. 23 (7/12): 172; TS: Papilio altheae Hübner
Lavatheria Verity, 1940; Le Farfalle diurn. d'Italia 1: 11, 22; TS: Papilio lavatherae Esper

Carcharodus; [LepIndex]; [CoL] ; [iNaturalist] ; [NHM-BMW]
Carcharodus (Pyrginae); [AfrL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 8
Lavatheria; [NHM-BMW]

Carcharodus alceae (Esper, 1780)

SEU, CEU, Temperate Asia (S.Siberia - Arabia, N.India). See [maps] Warcabnik ślazowiec Piquitos castana Mallow Skipper

Papilio alceae Esper, 1780; Die Schmett. Th. I, Bd. 2 (1): 4, pl. 51, f. 3; TL: Erlangen, Germany
malvarum (Hoffmannsegg, 1804) (Papilio); Mag. f. Insektenk. 3: 193, 203
Hesperia malvarum var. australis Zeller, 1847; Isis von Oken 1847 (4): 286
Hesperia malvarum var. nostras Zeller, 1847; Isis von Oken 1847 (4): 286

Carcharodus alceae alceae; [iNaturalist]
Carcharodus alceae gooraisa; [LepIndex]; [CoL]
Carcharodus alceae insolatrix; [LepIndex]; [CoL]
Carcharodus alceae; [H&R], 328; [BOW]: pl. 11, f. 11; [BRU]: 108, pl. 2, f. 1-3; [BMAT]: 106 (all N.African records are Carcharodus tripolina); Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 9; [AfrL]; [Otakar Kudrna]; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [SVF]
Spilothyrus alceae; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 506
Spilothyrus alceae var. australis; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 506

600x476 (~104Kb) Italy, Ravenna, 11 May 2006, Photo © Maurizio Elia

500x324 (~47Kb) Turunc, SW Turkey 14/04/95, Photo © Simon Coombes

C. a. wissmanni Warnecke, 1934

Carcharodus alceae wissmanni; [AfrL]; [LepIndex]; [CoL] ; [iNaturalist]

C. a. swinhoei Watson, 1893 Afghanistan, NW.India, ...

Carcharodus swinhoei Watson, 1893; Proc. Zool. Soc. Lond. 1893 (1): 68

Carcharodus alceae swinhoei; [LepIndex]; [CoL] ; [iNaturalist]

Carcharodus tripolina (Verity, 1925)

SWEU, N.Africa. See [maps]

Erynnis alceae tripolina Verity, 1925; Ent. Rec. J. Var. 37: 54; TL: Garian plateau (Libya)

Carcharodus alceae tripolina; [LepIndex]; [CoL]
Carcharodus tripolina; [BMAT]: 94, pl. 31, f. 6-10; [iNaturalist] ; [LEPIFORUM]

Carcharodus lavatherae (Esper, 1783)

NAF, SEU, Asia Minor - Iran. See [maps] Piquitos clara Marbled Skipper Ziestfalter

Papilio lavatherae Esper, 1783; Die Schmett. Th. I, Bd. 2 (7): 148, pl. 82, f. 4; TL: France & Switzerland

Carcharodus lavatherae internirufus; [LepIndex]; [CoL]
Carcharodus lavatherae tauricus; [LepIndex]; [CoL]
Carcharodus lavatherae; [BOW]: pl. 11, f. 11 (name); [H&R], 328; [BRU]: 108, pl. 2, f. 4-6; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 8; [LepIndex] ; [CoL] ; [MBE]

C. l. rufescens Oberthür, 1915 Morocco (Middle Atlas), Algeria, Tunisia?

Carcharodus lavaterae[sic] var. rufescens Oberthür, 1915; Étud. Lépid. Comp. 10: 407, pl. 296, f. 4433; TL: Sebdou (Algeria)
Carcharodus lavaterae[sic] internicus Rothschild, 1914; Novit. Zool. 21: 311; TL: Aïn Sefra, Guelt-es-Stel (Algeria)
Erynnis lavatherae australissima Verity, 1925; Ent. Rec. J. Var. 37: 41; TL: Sebdou, Lambessa (Algeria)

Carcharodus lavatherae rufescens; [BMAT]: 95, pl. 31, f. 11-15

Carcharodus baeticus (Rambur, 1840)

SWEU, it, Asia Minor, Iran. See [maps] Southern Marbled Skipper Piquitos

Syrichtus baeticus Rambur, 1840; Faune ent. Andal. 2 (4): pl. 12, f. 3-4
marrubii Rambur, 1840
- = Carcharodus boeticus boeticus; [H&R], 329
Spilothyrus marrubii fonti de Sagarra, 1930; Butll. Inst. catal. Hist. nat. (2) 10 (7): 118

Carcharodus baeticus; [Otakar Kudrna]; [MBE]
Carcharodus boeticus boeticus[sic, baeticus]; [H&R], 329
Carcharodus boeticus; [LepIndex] ; [CoL]
Carcharodus boeticus[sic, baeticus]; [H&R], 329; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 9
Spilothyrus altheae var. baeticus; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 506

Subgenus Reverdinus

Reverdinus; [NHM-BMW]

Carcharodus floccifera (Zeller, 1847)

SEU, Bulgaria, S.Siberia, Asia Minor. See [maps] Warcabnik szantawiec Piquitos serrana Tufted Skipper

Hesperia floccifera Zeller, 1847; Isis von Oken 1847 (4): 286; TL: Syracuse, Sicily
Papilio altheae Hübner, [1800-1803]; Samml. eur. Schmett. [1]: pl. 90, f. 452-453 (preocc. Papilio altheae Esper, [1783])
- = Carcharodus flocciferus; [H&R], 330
Carcharodus imperator Hemming, 1934; Stylops 3 (5): 99 (repl. Papilio altheae Hübner, [1800-1803])
alchymillae Hemming, 1936
- = Carcharodus flocciferus; [H&R], 330
gemina Lederer, 1852

Carcharodus floccifera; [BMAT]; [LepIndex] ; [CoL]
Carcharodus flocciferus; [H&R], 330; [BRU]: 109, pl. 2, f. 7-9; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 9; Yakovlev, 2012, Nota lepid. 35 (1): 61; [MBE]

533x400 (~40Kb) upperside An open pine forest on a high left bank of the Berd' River 1 km upstream of the village Starososedovo. Iskitim district, Novosibirsk Province, West Siberia, Russia. 2nd July 1994, Photo © Oleg Kosterin	800x800 (~133Kb) upperside An open pine forest on a high left bank of the Berd' River 1 km upstream of the village Starososedovo. Iskitim district, Novosibirsk Province, West Siberia, Russia. 2nd July 1994, Photo © Oleg Kosterin	604x674 (~123Kb) upperside Russia, Moscow area, 10.7.2019 (36°25'E, 56°00'N), Photo © D. Smirnov
644x584 (~110Kb) underside Russia, Moscow area, 10.7.2019 (36°25'E, 56°00'N), Photo © D. Smirnov

C. f. habiba (Kauffmann, 1955) Morocco (W.Rif Mts)

Reverdinus floccifer habiba Kauffmann, 1955; Mitt. schweiz. ent. Ges. 28 (3): 288; TL: Bab el Taza (Morocco)

Carcharodus floccifera habiba; [BMAT]: 95, pl. 31, f. 16-18; [LepIndex]; [CoL]

Carcharodus orientalis Reverdin, 1913

Montenegro, Greece, Asia Minor, Palestine, N.Iran. See [maps] Oriental Skipper

Carcharodus orientalis Reverdin, 1913; Bull. Soc. Lép. Genève, 2 (4): 230, f. 4, pl. 21, f. 14; TL: Peloponnesos, Greece

Carcharodus floccifera orientalis; [LepIndex]; [CoL]
Carcharodus orientalis; [H&R], 330; [BRU], 109; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 9; [MBE]

C. o. orientalis

Carcharodus orientalis orientalis; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 9

C. o. teberdininus Devyatkin, 1990 Caucasus Major

orientalis teberdininus Devyatkin, 1990; Ent. Obozr. 69 (4): 934

Carchadorus orientalis teberdininus; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 9

Carcharodus stauderi Reverdin, 1913

Morocco - Cyrenaica, Greece, Asia Minor - Palestine, N.Iran. See [maps]

Carcharodus stauderi Reverdin, 1913; Bull. Soc. Lép. Genève, 2 (4): 225, pl. 21, f. 5, 12; TL: El Kantara (Algeria)

Carcharodus boeticus[sic] stauderi; [H&R], 330
Carcharodus stauderi barcaeus; [LepIndex]; [CoL]
Carcharodus stauderi ramses; [LepIndex]; [CoL]
Carcharodus stauderi; [BRU]: 109, pl. 2, f. 13-15; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 9; [LepIndex] ; [CoL]

C. s. stauderi N.Africa

Erynnis stauderi f. fulvissima Verity, 1925; Ent. Rec. J. Var. 37: 43; TL: Algeria
Erynnis stauderi f. obscura Verity, 1925; Ent. Rec. J. Var. 37: 44; TL: Algeria
Carcharodus stauderi romeii Rothschild, 1933; Novit. Zool. 38: 323; TL: Ketama (Morocco)
Reverdinus stauderi f. rungsi Picard, 1950; Bull. Soc. Sci. nat. Maroc 28: 118-119; TL: Ifrane (Morocco)

Carcharodus stauderi romeii; [LepIndex]; [CoL]
Carcharodus stauderi stauderi; [BMAT]: 95, pl. 31, f. 19-20

C. s. ambigua Verity, 1925 S.Transcaucasia

Carcharodus stauderi ambigua; [LepIndex]; [CoL]

Carcharodus dravira (Moore, [1875])

N.Iran - Pakistan, N.India. See [maps]

Pyrgus dravira Moore, [1875]; Proc. zool. Soc. Lond. 1874 (4): 576, pl. 67, f. 5; TL: Kashmir
balucha Evans, 1932
Carcharodus dentatus Alberti, 1940; Mitt. Münch. Ent. Ges. 30: 236, pl. 2, f. 5d; TL: Badachschan, Sebaktal, Alpenwiesenzone, 2800-3000m

Carcharodus dravira; [BRU]: 110, pl. 2, f. 16-18; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 9
Carcharodus floccifera dravira; [LepIndex]; [CoL]
Pyrgus dravira; de Nicéville, [1884], J. Asiat. Soc. Bengal 52 Pt.II (2/4): 88, pl. 10, f. 5 ♀

References:

[AfrL] Mark C. Williams
Checklist of Afrotropical Papilionoidea and Hesperoidea; Compiled by Mark C. Williams, 7th ed. (2008) (April 2007); Afrototropical Butterflies; e-mail: Mark C. Williams
[CoL] Catalogue of Life
Bánki, O., Roskov, Y., Döring, M., Ower, G., Hernández Robles, D. R., Plata Corredor, C. A., Stjernegaard Jeppesen, T., Örn, A., Vandepitte, L., Hobern, D., Schalk, P., DeWalt, R. E., Ma, K., Miller, J., Orrell, T., Aalbu, R., Abbott, J., Adlard, R., Aedo, C., et al. (2024). Catalogue of Life Checklist (Version 2024-03-26); https://doi.org/10.48580/dfz8d
[D. Smirnov]
Russia, Moscow; e-mail: D. Smirnov
[ESP] :ESP#c Heppner, 1981
[EVERSMANNIA]
Eversmannia contents
[HBN] Francis Hemming, 1937
HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[Heiner Ziegler]
Switzerland / Schweiz; www.pieris.ch
[LEPIFORUM]
Lepiforum; Bestimmung von Schmetterlingen (Lepidoptera) und ihren Präimaginalstadien
[LepIndex] Global Lepidoptera Index
Beccaloni, G., Scoble, M., Kitching, I., Simonsen, T., Robinson, G., Pitkin, B., Hine, A., Lyal, C., Ollerenshaw, J., Wing, P., & Hobern, D. (2024). Global Lepidoptera Index. In O. Bánki, Y. Roskov, M. Döring, G. Ower, D. R. Hernández Robles, C. A. Plata Corredor, T. Stjernegaard Jeppesen, A. Örn, L. Vandepitte, D. Hobern, P. Schalk, R. E. DeWalt, K. Ma, J. Miller, T. Orrell, R. Aalbu, J. Abbott, R. Adlard, C. Aedo, et al., Catalogue of Life Checklist (1.1.24.106); https://doi.org/10.48580/dg4lg-49xk
[MBE] Moths and Butteflies of Europe
Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[Maurizio Elia]
Ravenna, Italy; e-mail: Maurizio Elia
[NHM-BMW] Natural History Museum
Butterflies and Moths of the World; Generic Names and their Type-species
[Oleg Kosterin] Institue of Cytology & Genetics
Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[SEG]
Schweizerische Entomologische Gesellschaft; Mitteilungen der Schweizerischen Entomologischen Gesellschaft; = Bulletin de la Société Entomologique Suisse = Journal of the Swiss Entomological Society
[SVF]
Svenska fjärilar; Naturhistoriska riksmuseet; (Sweden)
[Simon Coombes]
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[iNaturalist]
iNaturalist
[maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.

Some related literature:

Alberti, 1940
Über einige neue oder wenig bekannte palaearktische Hesperiiden nebst Revision der Hesperia staudingeri-Gruppe Mitt. Münch. Ent. Ges. 30 : 235-254, 10fgs
de Nicéville, [1884]
On new and little known Rhopalocera from the Indian Region J. Asiat. Soc. Bengal 52 Pt.II (2/4) : 65-91, pl. 1, 9-10
Esper, 1780; Esper, 1783
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Fortsetzung. Band 2 Die Schmett. Th. I, Bd. 2 (1): 1-36, pl. 51-56 (1780), (2): 37-68, pl. 57-62 (1781), (3): 69-100, pl. 63-68 (1781), (4): 101-124, pl. 69-74 (1781), (5): 125-140, pl. 75-80 (1782), (7): 141-156, pl. 81-82 (1783), (8): 157-172, pl. 83-88 (1783), (9): 173-184, pl. 89-92 (1784), (28): 185-190, pl. 93 (1786)
Grum-Grshimailo, 1890
Le Pamir et sa faune lépidoptérologique in Romanoff, Mém. Lép. 4 : 1-576, pl. A,1-21
Hemming, 1934
Some notes in the nomenclature of Palaearctic and African Rhopalocera Stylops 3 (5) : 97-99
[H&R]; Higgins & Riley, 1970
A Field Guide to the Butterflies of Britain and Europe
Hoffmannsegg, 1804
Alphabetisches Verzeichniss zu J. Hübner's Abbildungen der Papilionen mit den beigefügten vorzüglichsten Synonymen Mag. f. Insektenk. 3 : 181-206
Hübner, [1800-1803]
Sammlung europäischer Schmetterlinge. I. Papiliones - Falter ("Erste Band") Samml. eur. Schmett. [1] : 3-74, : pl. 1-88, f. 1-446 (1799-1800), : pl. 89-96, f. 447-490 (1800-1803), : pl. 97-114, f. 491-588 (1803-1804), : pl. 115-119, f. 589-612 (1805), : pl. 120-124, f. 613-636 (1805-1806), : pl. 125-128, f. 637-652 (1806-1808), : pl. 129-144, f. 653-727 (1808-1813), : pl. 145-147, f. 728-741 (1814-1816), : pl. 148-150, f. 742-756 (1816-1817), : pl. 151, f. 757-762 (1818-1819), : pl. 152-154, f. 757-774 (1819-1822), : pl. 155, f. 775-778 (1822-1823), : pl. 156-161, f. 779-806 (1823), : pl. 162-175, f. 807-874 (1823-1824), : pl. 176, f. 869-874 (1824-1825), : pl. 177-181, f. 875-895 (1825-1826), : pl. 182-187, f. 896-981 (1827-1828), : pl. 188-195, f. 982-968 (1828-1832), : pl. 196, f. 969-974 (1832-1833), : pl. 197-201, f. 975-997 (1834-1836), : pl. 202-203, f. 998-1009 (1836-1838), : pl. 204-207, f. 1010-1029 (1837-1838)
Hübner, [1819]
Verzeichniss bekannter Schmettlinge, 1816-[1826] Verz. bek. Schmett. (1): [1-3], 4-16 (1816), (2): 17-32 (1819), (3): 33-48 (1819), (4): 49-64 (1819), (5): 65-80 (1819), (6): 81-96 (1819), (7): 97-112 (1819), (8): 113-128 (1819), (9): 129-144 (1819), (10): 145-160 (1819), (11): 161-176 (1819), (12): 177-192 (1820), (13): 193-208 (1820), (14): 209-224 (1821), (15): 225-240 (1821), (16): 241-256 (1821), (17): 257-272 (1823), (18): 273-288 (1823), (19): 289-304 (1823), (20): 305-320 (1825), (21): 321-336 (1825), (22): 337-352 (1825), (23-27): 353-431 ([1825])
Kauffmann, 1955
Reverdinus floccifer habiba n. ssp. (Lépidopt. Hesperiidae) Mitt. schweiz. ent. Ges. 28 (3) : 288-290
Korb & Bolshakov, 2011
[A catalogue of butterflies (Lepidoptera: Papilioformes) of the former USSR. Second edition, reformatted and updated] (in Russian) Eversmannia Suppl. 2 : 1-123
[BOW]; Lewis, 1974
Butterflies of the World
Moore, [1875]
Descriptions of new Asiatic Lepidoptera Proc. zool. Soc. Lond. 1874 (4) : 565-579, pl. 66-67
Oberthür, 1915
Faune des Lépidoptères de la Barbarie, Algerie Étud. Lépid. Comp. 10 : 1-459, pl. 276-277,277bis,278-315
Ragusa, 1919
Elenco dei lepidotteri di Sicilia Natural. Sicil. 23 (7/12) : 144-178
Rambur, 1840
Faune entomologique de l'Andalusie. 1837-1840 (Vol. 2) Faune ent. Andal. 2 (1): pl. 14-15, 18 (1837), 2 (2): 1-16, pl. 1-4 (1838), 2 (3): 17-96, pl. 6-7, 9, 11 (1838), 2 (4): 97-176, pl. 5, 8, 10, 12, 17 (1839), 2 (5): 177-336 (1840)
Reverdin, 1913
Notes sur les genres Hesperia et Carcharodus Bull. Soc. Lép. Genève, 2 (4): 212-234, pl. 21-22
Rothschild, 1914
A preliminary account of the Lepidopterous fauna of Guelt-es-Stel, central Algeria Novit. Zool. 21: 299-357 (1914), 22 (2): 186-191 (1915)
Rothschild, 1933
On a collection of Lepidoptera from Spanish Morocco Novit. Zool. 38 : 315-330
de Sagarra, 1930
Anotacions a la lepidopterologia Ibérica V (2). Formes noves de lepidòpters ibérics Butll. Inst. catal. Hist. nat. (2) 10 (7) : 110-118
[BMAT]; Tennent, 1996
The Butterflies of Morocco, Algeria and Tunisia : 1-217
[BRU]; Tuzov, Bogdanov, Devyatkin, Kaabak, Korolev, Murzin, Samodurov, Tarasov, 1997
Guide to the Butterflies of Russia and adjacent territories: Hesperiidae, Papilionidae, Pieridae, Satyridae Butts. Russia adj. terr. 1
Verity, 1925
New races and forms of Palaearctic Grypocera Ent. Rec. J. Var. 37 : 41-44,54-57,72-76
Watson, 1893
A proposed classification of the Hesperiidae, with a revision of the genera Proc. Zool. Soc. Lond. 1893 (1) : 3-133, pl. 1-3
Yakovlev, 2012
Checklist of Butterflies (Papilionoidea) of the Mongolian Altai Mountains, including descriptions of new taxa Nota lepid. 35 (1) : 51-96
Zeller, 1847
Bemerkungen über die auf einer Reise Nach Italien und Sicilien Beobachteten Schmetterlingsarten Isis von Oken 1847 (2): 121-159, (3): 213-233, (4): 284-308, (6): 401-457, (7): 481-522, (8): 561-594, (9): 641-673, (10): 721-771, (11): 801-859, (12): 881-914

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