Chlosyne

Chlosyne Butler, 1870

Chlosyne Butler, 1870; Cistula ent. 1 (3): 38 (repl. Synchloe Doubleday, 1844); TS: Papilio janais Drury
Morpheis Geyer, [1833]; in Hübner, Samml. exot. Schmett. 3: pl. [5] (preocc. Morpheis Hübner, [1820]); TS: Morpheis ehrenbergii Geyer
- = Chlosyne; [NL4A], 252; Pelham, 2008, J. Res. Lepid. 40: 371
Synchloe Doubleday, 1844; List. lepid. Ins. Brit. Mus. 1: 76 (preocc. Synchloe Hübner, 1818); TS: Papilio janais Drury
- = Chlosyne; [NL4A], 252; Pelham, 2008, J. Res. Lepid. 40: 371
Coatlantona Kirby, 1871; Syn. Cat. diurn. Lep.: 178 (repl. Synchloe Doubleday, 1844); TS: Papilio janais Drury
- = Chlosyne; [NL4A], 252; Pelham, 2008, J. Res. Lepid. 40: 371
Anemeca Kirby, 1871; Syn. Cat. diurn. Lep.: 179 (repl. Morpheis Geyer, [1833]); TS: Morpheis ehrenbergii Geyer
- = Chlosyne; [NL4A], 252; Pelham, 2008, J. Res. Lepid. 40: 371
Charidryas Scudder, 1872; 4th Ann. Rep. Peabody Acad. Sci. (1871): 47; TS: Melitaea nycteis Doubleday
- = Chlosyne; [NL4A], 252; Pelham, 2008, J. Res. Lepid. 40: 371; [Wahlberg, 2023]
Limnaecia Scudder, 1872; 4th Ann. Rep. Peabody Acad. Sci. (1871): 47 (preocc. Limnaecia Stainton, 1851); TS: Melitaea harrisii Scudder
- = Charidryas;
- = Chlosyne; [NL4A], 252; Pelham, 2008, J. Res. Lepid. 40: 372; [Wahlberg, 2023]
Thessalia Scudder, 1875; Bull. Buffalo Soc. nat. Sci. 2: 238, 265; TS: Melitaea leanira C. & R. Felder
- = Chlosyne; [NL4A], 252; Pelham, 2008, J. Res. Lepid. 40: 372; [Wahlberg, 2023]

Anemeca; [LepIndex]; [CoL] ; [CoL] ; [NHM-BMW]
Charidryas; [NHM-BMW]
Chlosyne; [P&N23]; [LepIndex]; [CoL] ; [iNaturalist] ; [NHM-BMW]
Chlosyne (Melitaeina); Zhang, Cong, Shen, Opler & Grishin, 2021, Taxonomic Rep. 9 (3): 42
Chlosyne (Melitaeini); Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (included Charidryas also); [Wahlberg, 2023]
Chlosyne ([Chlosynina]); Pelham, 2008, J. Res. Lepid. 40: 371
Chlosyne ([I]); [NL4A], 252
Coatlantona; [NHM-BMW]
Morpheis; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 211
Synchloe; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 176; [NHM-BMW]
Thessalia; [LepIndex]; [CoL] ; [NHM-BMW]
Thessalia (Melitaeini); Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167

700x605 (~70Kb) upperside USA: Bridge Creek Trailhead, North Cascades Highway, (48°30'30N 120°43'00W), Chelan Co., WA, 28.7.1999, Photo © Markku Savela	700x679 (~62Kb) underside USA: Bridge Creek Trailhead, North Cascades Highway, (48°30'30N 120°43'00W), Chelan Co., WA, 28.7.1999, Photo © Markku Savela	550x619 (~38Kb) underside USA: Table Mtn, Kittitas Co., WA, 30.7.1999, Photo © Markku Savela
600x829 (~63Kb) upperside USA: nr Crystal Mountain / FR9726 (47°14'37N 120°38'39W), Kittitas Co., WA, 11.7.2000, Photo © Markku Savela	600x722 (~55Kb) underside USA: nr Crystal Mountain / FR9726 (47°14'37N 120°38'39W), Kittitas Co., WA, 11.7.2000, Photo © Markku Savela

The lacinia species-group

Chlosyne janais (Drury, 1782)

Texas, Mexico - Colombia. See [maps] Janais Patch

Papilio janais Drury, 1782; Illust. Nat. Hist. Exot. Insects 3: index, 22, pl. 17, f. 5-6; TL: ["Brazil, Rio de Janeiro"]

Chlosyne janais; Dyar, 1903, Bull. U.S. nat. Mus. 52: 21; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name); [NACL], #4503; [NAB], 592; [BCR], 199; [EBW]; [BOW]: pl. 13, f. 13; [Opler]; [NL4A], #2083; Pelham, 2008, J. Res. Lepid. 40: 372; [MPG] ; [P&N23]; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [BOA] ; [BAMONA]
Synchloe janais; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 178; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 671

C. j. janais

Chlosyne janais janais; [NL4A], #2083a; Pelham, 2008, J. Res. Lepid. 40: 372; [P&N23]; [iNaturalist] ; [BOA]

C. j. gloriosa Bauer, [1960] Mexico

Chlosyne gloriosa Bauer, [1960]; J. Lep. Soc. 13: 165; TL: Tepic, Nayarit, Mexico

Chlosyne gloriosa; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name)
Chlosyne janais gloriosa; [NL4A], #2083b; [LepIndex]; [CoL] ; [iNaturalist] ; [BOA]

C. j. marianna Röber, 1914 Mexico

Chlosyne hyperia marianna Röber, 1914; in Seitz, Gross-Schmetterl. Erde 5: 452; TL: Mexico
Chlosyne hyperia var. irrubescens Hall, 1917; Entomologist 50: 163; TL: Mexico, Morelos
- = ; [NL4A], #2083c

Chlosyne janais irrubescens; [LepIndex]; [CoL]
Chlosyne janais marianna; [NL4A], #2083c; [LepIndex]; [CoL] ; [iNaturalist] ; [BOA]
Chlosyne marianna; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name)

Chlosyne rosita Hall, 1924

-. See [maps] Rosita Patch

Chlosyne rosita Hall, 1924; Entomologist 57: 241; TL: Guatemala

Chlosyne rosita; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name); [NACL], #4504; [NAB], 593 (note); [Opler]; [NL4A], #2091; Pelham, 2008, J. Res. Lepid. 40: 373; [MPG] ; [P&N23]; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [BOA] ; [BAMONA]

C. r. rosita Guatemala

Chlosyne rosita rosita; [NL4A], #2091a; [iNaturalist] ; [BOA]

C. r. montana Hall, 1924 Mexico

Chlosyne lacinia montana Hall, 1924; Entomologist 57: 241; TL: Mexico, Mt Popacatapetl, 10000ft

Chlosyne lacinia montana; [LepIndex]; [CoL]
Chlosyne montana;
Chlosyne rosita montana; [NL4A], #2091d; Pelham, 2008, J. Res. Lepid. 40: 373; [P&N23]; [iNaturalist] ; [BOA]

C. r. browni Bauer, [1961] Mexico

Chlosyne rosita browni Bauer, [1961]; J. Lep. Soc. 14 (2): 150; TL: El Salto, 1600ft, San Luis Potosí, Mexico

Chlosyne rosita browni; [NACL], #4504a; [NL4A], #2091b; Pelham, 2008, J. Res. Lepid. 40: 373; [P&N23]; [LepIndex]; [CoL] ; [iNaturalist] ; [BOA]

C. r. riobalsensis Bauer, [1961] Mexico

Chlosyne riobalsensis Bauer, [1961]; J. Lep. Soc. 14 (2): 152; TL: Mexcala, 2000ft, Guerrero, Mexico

Chlosyne riobalsensis; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name); [LepIndex] ; [CoL]
Chlosyne rosita riobalsensis; [NL4A], #2091e; [iNaturalist] ; [BOA]

C. r. mazarum Miller & Rotger, 1979 Mexico

Chlosyne mazarum Miller & Rotger, 1979; Bull. Allyn Mus. 54: 3; TL: Mexico, Morelos, Paraje, Rancho Viejo

Chlosyne mazarum; [LepIndex] ; [CoL]
Chlosyne rosita mazarum; [NL4A], #2091c; [iNaturalist]

Chlosyne hippodrome (Geyer, 1837)

Mexico - Colombia. See [maps]

Araschnia hippodrome Geyer, 1837; in Hübner, Zuträge. Samml. exot. Schmett. 5: f. 863-864; TL: Mexico

Chlosyne hippodrome; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name); [BCR], 199; [NL4A], #2082; [LepIndex] ; [CoL] ; [iNaturalist] ; [BOA] ; [BAMONA]

C. h. hippodrome Mexico

Chlosyne hippodrome hippodrome; [NL4A], #2082a; [iNaturalist] ; [BOA]

C. h. fabricii Higgins, 1960

hippodrome fabricii Higgins, 1960; (repl. name); TL: "Indiis"
Papilio hyperia Fabricius, 1793; Ent. Syst. 3 (1): 119 (preocc. Cramer, 1779); TL: "Indiis"
- = ; [NL4A], #2082b

Chlosyne hippodrome fabricii; [NL4A], #2082b; [LepIndex]; [CoL]
Synchloe hyperia; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 179; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 672

Chlosyne gaudialis (Bates, 1864)

Panama, .... See [maps]

Synchloe gaudialis Bates, 1864; Ent. mon. Mag. 1 (4): 84, n. 40; TL: Guatemala

Chlosyne gaudealis[sic?]; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name)
Chlosyne gaudealis[sic]; [BCR], 200 (missp.)
Chlosyne gaudialis; [EBW]; [NL4A], #2080; [LepIndex] ; [CoL] ; [iNaturalist] ; [BOA] ; [BAMONA]
Synchloe gaudialis; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 182; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 673
?Coatlantona gaudialis;

681x619 (~46Kb) Belize, Central America, Photo © John Shuey

C. g. gaudialis Guatemala, Nicaragua

Chlosyne gaudialis laeta Röber, 1914; in Seitz, Gross-Schmetterl. Erde 5: 452; TL: Nicaragua
- = ; [NL4A], #2080a

Chlosyne gaudialis gaudialis; [NL4A], #2080a; [iNaturalist] ; [BOA]
Chlosyne gaudialis laeta; [LepIndex]; [CoL]

C. g. wellingi Miller & Rotger, 1979 Mexico

Chlosyne gaudialis wellingi Miller & Rotger, 1979; Bull. Allyn Mus. 54: 1; TL: Mexico, Oaxaca, Candelaria Loxicha, 500m

Chlosyne gaudialis wellingi; [NL4A], #2080b; [LepIndex]; [CoL] ; [iNaturalist] ; [BOA]

Chlosyne narva (Fabricius, 1793)

Nicaragua - Venezuela. See [maps]

Papilio narva Fabricius, 1793; Ent. Syst. 3 (1): 249; TL: "Africa"

Chlosyne narva; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name); [BCR], 200; [BOW]: pl. 34, f. 7; [NL4A], #2089; [LepIndex] ; [CoL] ; [iNaturalist] ; [BOA]

Misidentified/misapplied:

?Synchloe narva (= Chlosyne narva bonpland); Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 3: pl. 20, f. 7-8

C. n. narva

Chlosyne narva narva; [NL4A], #2089a; [iNaturalist] ; [BOA]

C. n. bonpland (Latreille, [1809]) Panama

Cethosia bonpland Latreille, [1809]; in Humboldt & Bonpland, Voy. rég. équinoxiales..., 1 (4): 199, pl. 18, f. 5; TL: "[Ecuador]"
Chlosyne narva var. brunhilda Staudinger, 1885; in Staudinger & Schatz, Exot. Schmett. 1 (10): 96; TL: Panama
- = ; [NL4A], #2089b

Chlosyne narva bonpland; [NL4A], #2089b; [LepIndex]; [CoL] ; [iNaturalist] ; [BOA]
Synchloe bonplandi[sic, bonpland]; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 182; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 673

Chlosyne ehrenbergii (Geyer, [1833])

Mexico. See [maps]

Morpheis ehrenbergii Geyer, [1833]; in Hübner, Samml. exot. Schmett. 3: pl. [5]; TL: [Mexico]

Anemeca ehrenbergi;
Chlosyne ehrenbergi; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name); [BOW]: pl. 34, f. 18; [BAMONA]
Chlosyne ehrenbergii; [NL4A], #2075; [P&N23]; [LepIndex] ; [CoL] ; [iNaturalist] ; [BOA]
Morpheis ehrenbergi; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 211; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 682

Chlosyne lacinia (Geyer, 1837)

-. See [maps] Bordered Patch

Araschnia lacinia Geyer, 1837; in Hübner, Zuträge. Samml. exot. Schmett. 5: 25, f. 899-900; TL: Mexico
Synchloe nigrescens Wright, 1905; Butts. West Coast US: 170, pl. 21, f. 210

Chlosyne lacina[sic]; [EBW]
Chlosyne lacinia nigrescens; [LepIndex]; [CoL]
Chlosyne lacinia; Dyar, 1903, Bull. U.S. nat. Mus. 52: 21; [NACL], #4499; [NAB], 590; [BCR], 201; [BOW]: pl. 13, f. 14, pl. 34, f. 19; [Opler]; [NL4A], #2085; Pelham, 2008, J. Res. Lepid. 40: 377; [MPG] ; [P&N23]; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [BOA] ; [BAMONA]
Synchloe lacinia; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 176, 3: pl. 19, f. 6-17; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 671

C. l. lacinia Mexico, Guatemala, Panama, Costa Rica

Synchloe tellias Bates, 1864; Ent. mon. Mag. 1 (4): 84; TL: Guatemala
- = Chlosyne lacinia; Pelham, 2008, J. Res. Lepid. 40: 377
- = ; [NL4A], #2085a
Synchloe quehtala Reakirt, 1866; Proc. Acad. nat. Sci. Philad. 18 (3): 248; TL: Mexico
- = Chlosyne lacinia; Dyar, 1903, Bull. U.S. nat. Mus. 52: 21; Pelham, 2008, J. Res. Lepid. 40: 377
- = ; [NL4A], #2085a
Synchloe ardema Reakirt, [1867]; Proc. Acad. nat. Sci. Philad. 18 (4): 336; TL: Mexico, Veracruz
- = Chlosyne lacinia; Dyar, 1903, Bull. U.S. nat. Mus. 52: 21; Pelham, 2008, J. Res. Lepid. 40: 377
- = ; [NL4A], #2085a
Synchloe misera R. Felder, 1869; Verh. zool.-bot. Ges. Wien. 19: 471; TL: Mexico, Oaxaca; Morelos
- = Chlosyne lacinia; Dyar, 1903, Bull. U.S. nat. Mus. 52: 21; Pelham, 2008, J. Res. Lepid. 40: 377
- = ; [NL4A], #2085a
Synchloe pretona Boisduval, 1870; Considérations Lépid. Guatemala: 37; TL: Guatemala; Mexico
- = Chlosyne lacinia; Dyar, 1903, Bull. U.S. nat. Mus. 52: 21; Pelham, 2008, J. Res. Lepid. 40: 377
- = ; [NL4A], #2085a
Synchloë adelina Staudinger, 1876; Verh. zool.-bot. Ges. Wien, 25 (1): 102; TL: Panama
- = ; [NL4A], #2085a
- = Chlosyne lacinia; Pelham, 2008, J. Res. Lepid. 40: 377
Chlosyne lacinia lacinia f. indigens Higgins, 1960; Trans. R. ent. Soc. Lond. 112 (14): 408; TL: Costa Rica
- = ; [NL4A], #2085a
- = Chlosyne lacinia; Pelham, 2008, J. Res. Lepid. 40: 377

Chlosyne lacinia adelina; Dyar, 1903, Bull. U.S. nat. Mus. 52: 21; [LepIndex]; [CoL]
Chlosyne lacinia indigens; [LepIndex]; [CoL]
Chlosyne lacinia lacinia; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name); [NL4A], #2085a; [iNaturalist] ; [BOA]
Chlosyne lacinia quehtala; [LepIndex]; [CoL]
pretoria[sic]; Dyar, 1903, Bull. U.S. nat. Mus. 52: 21 (missp.)

C. l. saundersi (Doubleday, [1847]) Venezuela, Colombia, Ecuador, Bolivia, Argentina

Synchloe saundersi Doubleday, [1847]; Gen. diurn. Lep. (1): 186, (1): pl. 24, f. 2; TL: Venezuela
Synchloë mediatrix C. & R. Felder, [1867]; Reise Fregatte Novara, Bd 2 (Abth. 2) (3): 395; TL: Colombia
- = ; [NL4A], #2085d
Synchloë paupera C. & R. Felder, [1867]; Reise Fregatte Novara, Bd 2 (Abth. 2) (3): 395; TL: Colombia
- = Chlosyne lacinia; Dyar, 1903, Bull. U.S. nat. Mus. 52: 21
- = ; [NL4A], #2085d
Chlosyne mediatrix f. fassli Röber, [1914]; ; TL: Colombia
- = ; [NL4A], #2085d
Chlosyne mediatrix f. felderi Röber, 1914; in Seitz, Gross-Schmetterl. Erde 5: 451; TL: Colombia
- = ; [NL4A], #2085d
Chlosyne mediatrix f. fruhtorferi Röber, 1914; in Seitz, Gross-Schmetterl. Erde 5: 451; TL: Bolivia
- = ; [NL4A], #2085d
Chlosyne saundersi f. splendida Hayward, 1931; Rev. Soc. ent. argent. 4: 66, 181; TL: Argentina
- = ; [NL4A], #2085d
Chlosyne saundersi f. splendida ab. reducta Hayward, 1931; Rev. Soc. ent. argent. 4: 66, 181; TL: Argentina
- = ; [NL4A], #2085d
Chlosyne saundersi f. infuscata Hayward, 1935; Rev. Soc. ent. argent. 7: 186; TL: Argentina
- = ; [NL4A], #2085d

Chlosyne lacinia felderi; [LepIndex]; [CoL]
Chlosyne lacinia paupera; [LepIndex]; [CoL]
Chlosyne lacinia saundersi; Dyar, 1903, Bull. U.S. nat. Mus. 52: 21; Brown & Mielke, 1967, J. Lep. Soc. 21 (2) (2): 98; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name); [EcuL]; [NL4A], #2085d; [LepIndex]; [CoL] ; [iNaturalist] ; [BOA]
Synchloë saundersii; Doubleday, [1847], (missp.)

C. l. crocale (Edwards, 1874) Arizona, New Mexico, Mexico

Synchloe crocale Edwards, 1874; Trans. amer. ent. Soc. 5 (1): 17; TL: Arizona
Synchloe lacinia f. rufa Cockerell, 1893; Can. Ent. 25 (11): 286; TL: New Mexico
- = ; [NL4A], #2085c (part.); Pelham, 2008, J. Res. Lepid. 40: 377
Synchloe lacinia f. bicolor Cockerell, 1893; Can. Ent. 25 (11): 287; TL: New Mexico
- = ; [NL4A], #2085c (part.); Pelham, 2008, J. Res. Lepid. 40: 377
Synchloe lacinia f. nigra Cockerell, 1893; Can. Ent. 25 (11): 287; TL: New Mexico
- = ; [NL4A], #2085c (part.); Pelham, 2008, J. Res. Lepid. 40: 378
Synchloe lacinia ab. nigrescens Edwards, 1893; Can. Ent. 25 (11): 287; TL: New Mexico
- = ; [NL4A], #2085c; Pelham, 2008, J. Res. Lepid. 40: 378
Synchloe lacinia ab. rufescens Edwards, 1893; Can. Ent. 25 (11): 287; TL: Mexico, Chihuahua
- = ; [NL4A], #2085c; Pelham, 2008, J. Res. Lepid. 40: 377
Synchloe lacinia ab. rufescens Cockerell, 1894; Can. Ent. 26 (1): 30
- = ; [NL4A], #2085c
Chlosyne lacinia rufescens tr.f. inghami Gunder, 1928; Can. Ent. 60 (7): 167; TL: Arizona
- = Chlosyne lacinia adjutrix; [NL4A], #2085b
- = ; Pelham, 2008, J. Res. Lepid. 40: 378
Chlosyne lacinia f. flavida Higgins, 1960; Trans. R. ent. Soc. Lond. 112 (14): 409; TL: Mexico, Chihuahua
- = ; [NL4A], #2085c; Pelham, 2008, J. Res. Lepid. 40: 378

Chlosyne lacinia crocale; [NL4A], #2085c; [NACL], #4499a; Pelham, 2008, J. Res. Lepid. 40: 377; [P&N23]; [LepIndex]; [CoL] ; [iNaturalist] ; [BOA]
Chlosyne lacinia flavida; [LepIndex]; [CoL]

C. l. adjutrix (Scudder, 1875) Texas, New Mexico, Arizona

Chlosyne adjutrix Scudder, 1875; Bull. Buffalo Soc. nat. Sci. 2: 269; TL: Texas
Synchloe lacinia f. nigra Cockerell, 1893; ; TL: New Mexico
- = ; [NL4A], #2085b (part.)
Synchloe lacinia f. bicolor Cockerell, 1893; ; TL: New Mexico
- = ; [NL4A], #2085b (part.)
Synchloe lacinia f. rufa Cockerell, 1893; ; TL: New Mexico
- = ; [NL4A], #2085b (part.)

Chlosyne lacinia adjutrix; [NACL], #4499b; [NL4A], #2085b; Pelham, 2008, J. Res. Lepid. 40: 378; [P&N23]; [iNaturalist] ; [BOA]

?C. l. fruhstorferi Röber, 1914 Bolivia

Chlosyne mediatrix fruhstorferi Röber, 1914; in Seitz, Gross-Schmetterl. Erde 5: 451

Chlosyne lacinia fruhstorferi; [LepIndex]; [CoL]

Chlosyne californica (Wright, 1905)

California. See [maps] California Patch

Synchloe californica Wright, 1905; Butts. West Coast US: 170; TL: California
Chlosyne california[sic] ab. ♀ chinoi Gunder, 1924; Ent. News 35 (5): 157; TL: California
- = ; [NL4A], #2072; Pelham, 2008, J. Res. Lepid. 40: 376

Chlosyne californica; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name); [NACL], #4498; [NAB], 589; [Opler]; [NL4A], #2072; Pelham, 2008, J. Res. Lepid. 40: 376; [MPG] ; [P&N23]; [iNaturalist] ; [BOA] ; [BAMONA]
Chlosyne lacinia californica; [LepIndex]; [CoL]

Chlosyne melanarge (Bates, 1864)

Mexico - Costa Rica, Guatemala. See [maps]

Synchloe melanarge Bates, 1864; Ent. mon. Mag. 1 (4): 85; TL: Guatemala
Chlosyne paupera Boisduval, 1870; Considérations Lépid. Guatemala: 38 (preocc. C. & R. Felder, 1867); TL: Guatemala
- = ; [NL4A], #2088

Chlosyne melanarge; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name); [BCR], 200; [EBW]; [BOW]: pl. 34, f. 22; [NL4A], #2088; [LepIndex] ; [CoL] ; [iNaturalist] ; [BOA] ; [BAMONA]
Synchloe melanarge; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 179, 3: pl. 20, f. 9-10; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 672

Chlosyne erodyle (Bates, 1864)

Mexico - Costa Rica. See [maps]

Synchloe erodyle Bates, 1864; Ent. mon. Mag. 1 (4): 84; TL: Guatemala

Chlosyne erodyle; Dyar, 1903, Bull. U.S. nat. Mus. 52: 21; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name); [NACL], #4502; [BCR], 201; [BOW]: pl. 34, f. 16; [NL4A], #2077; [P&N23]; [LepIndex] ; [CoL] ; [iNaturalist] ; [BOA] ; [BOA] ; [BAMONA]
Synchloe erodyle; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 180, 3: pl. 20, f. 3-4; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 672

C. e. erodyle Guatemala

Synchloe erodyle Doubleday, [1845]; (nom. nud.)
- = ; [NL4A], #2077a

Chlosyne erodyle erodyle; [NL4A], #2077a; [iNaturalist] ; [BOA]

C. e. poecile (C. & R. Felder, 1867) Costa Rica - Colombia, Venezuela

Synchloë poecile C. & R. Felder, [1867]; Reise Fregatte Novara, Bd 2 (Abth. 2) (3): 396; TL: Colombia
Synchloe poecile C. & R. Felder, 1867

Chlosyne erodyle poecile; [NL4A], #2077b
Chlosyne poecile; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name); [BCR], 201; [LepIndex] ; [CoL] ; [iNaturalist] ; [BOA] ; [BAMONA]
Synchloe poecile; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 180, 3: pl. 20, f. 1-2; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 672

C. e. rubrigutta Röber, 1914 Colombia

Chlosyne fasciata rubrigutta Röber, 1914; in Seitz, Gross-Schmetterl. Erde 5: 452; TL: Colombia

Chlosyne erodyle rubrigutta; [NL4A], #2077c
Chlosyne poecile rubrigutta; [LepIndex]; [CoL] ; [BOA]

ssp. Venezuela

Chlosyne ; Lamas, MS
Chlosyne erodyle ssp.; [NL4A], #2077d

Chlosyne melitaeoides (C. & R. Felder, [1867])

Mexico. See [maps]

Synchloe melitaeoides C. & R. Felder, [1867]; Reise Fregatte Novara, Bd 2 (Abth. 2) (3): 396; TL: Mexico

Chlosyne marina melitaeoides; [NL4A], #2087c; [LepIndex]; [CoL]
Chlosyne melitaeoides; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name); [Opler]; Pelham, 2008, J. Res. Lepid. 40: 372; de la Maza & de la Maza, 2019, Rev. Soc. Mex. Lep. (ns) 6 (2): 4; [MPG] ; [P&N23]; [iNaturalist] ; [BOA] ; [BAMONA]
Synchloe melitaeoides; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 181

C. m. melitaeoides S.Texas, Mexico

Chlosyne melitaeoides melitaeoides; de la Maza & de la Maza, 2019, Rev. Soc. Mex. Lep. (ns) 6 (2): 4; [iNaturalist]

C. m. haghenbecki de la Maza & de la Maza, 2019 Mexico (Oaxaca)

Chlosyne melitaeoides haghenbecki de la Maza & de la Maza, 2019; Rev. Soc. Mex. Lep. (ns) 6 (2): 4; TL: Oaxaca, Tepelmeme, Arroyo el Aguacate

Chlosyne melitaeoides haghenbecki; [iNaturalist]

Chlosyne eumeda (Godman & Salvin, 1894)

Mexico. See [maps]

Synchloë eumeda Godman & Salvin, 1894; Ann. Mag. nat. Hist. (6) 14 (80): 96; TL: Mexico, Guerrero
Synchloë dryope Godman & Salvin, 1894; Ann. Mag. nat. Hist. (6) 14 (80): 96; TL: Mexico, Jalisco
- = Chlosyne marina eumeda; [NL4A], #2087b
- = ; Pelham, 2008, J. Res. Lepid. 40: 373
Chlosyne fasciata Röber, 1914; in Seitz, Gross-Schmetterl. Erde 5: 452; TL: Mexico, Guerrero
- = Chlosyne marina eumeda; [NL4A], #2087b
- = ; Pelham, 2008, J. Res. Lepid. 40: 373

Chlosyne eumeda dryope; [LepIndex]; [CoL]
Chlosyne eumeda; [EBW]; [BOW]: pl. 34, f. 20; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 373; [MPG] ; [P&N23]; [LepIndex] ; [CoL] ; [iNaturalist] ; [BOA] ; [BOA] ; [BAMONA]
Chlosyne marina eumeda; [NL4A], #2087b
Synchloe dryope; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 672, 3: pl. 108, f. 3-4
Synchloe eumeda; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 672, 3: pl. 108, f. 1-2

Chlosyne marina (Geyer, 1837)

-. See [maps]

Araschnia marina Geyer, 1837; in Hübner, Zuträge. Samml. exot. Schmett. 5: 20, f. 877-878; TL: Mexico

Chlosyne marina; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name); [BOW]: pl. 34, f. 21; [NL4A], #2087; [LepIndex] ; [CoL] ; [iNaturalist] ; [BOA] ; [BOA] ; [BOA] ; [BAMONA]
Synchloe marina; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 181; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 673

C. m. marina Mexico

Synchloe hylaeus Godman & Salvin, 1894; Ann. Mag. nat. Hist. (6) 14 (80): 97; TL: Mexico, Durango
- = ; [NL4A], #2087a

Chlosyne marina hylaeus; [LepIndex]; [CoL]
Chlosyne marina marina; [NL4A], #2087a; [iNaturalist] ; [BOA]
Synchloe hylaeus; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 673

Chlosyne endeis (Godman & Salvin, 1894)

-. See [maps]

Synchloe endeis Godman & Salvin, 1894; Ann. Mag. nat. Hist. (6) 14 (80): 97; TL: Mexico, Nayarit

Chlosyne endeis; [NACL], #4501; [Opler]; [NL4A], #2076; Pelham, 2008, J. Res. Lepid. 40: 372; [MPG] ; [P&N23]; [LepIndex] ; [CoL] ; [iNaturalist] ; [BOA] ; [BAMONA]
Synchloe endeis; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 673, 3: pl. 108, f. 5-6

C. e. endeis Mexico

Chlosyne endeis endeis; [NL4A], #2076a; [iNaturalist]

C. e. pardelina Scott, 1986 Mexico

Chlosyne endeis pardelina Scott, 1986; Butts. N. America: 301
Chlosyne endeis f. pardelina Higgins, 1960; Trans. R. ent. Soc. Lond. 112 (14): 420; TL: Mexico, Coahuila
- = ; Pelham, 2008, J. Res. Lepid. 40: 372

Chlosyne endeis pardelina; [NL4A], #2076b; Pelham, 2008, J. Res. Lepid. 40: 372; [LepIndex]; [CoL] ; [iNaturalist]

Chlosyne definita (Aaron, [1885])

Texas. See [maps] Definite Patch

Melitaea definita Aaron, [1885]; Papilio 4 (9/10): 176; TL: Texas

Chlosyne definita; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name); [NACL], #4500; [NAB], 591; [Opler]; [NL4A], #2074; Pelham, 2008, J. Res. Lepid. 40: 372; [MPG] ; [P&N23]; [LepIndex] ; [CoL] ; [iNaturalist] ; [BOA] ; [BAMONA]
Thessalia definita; Dyar, 1903, Bull. U.S. nat. Mus. 52: 19

C. d. definita Texas, Mexico

Melitaea albiplaga Aaron, [1885]; Papilio 4 (9/10): 175; TL: Nueces Co., Texas
- = ; [NL4A], #2074a; Pelham, 2008, J. Res. Lepid. 40: 372
Melitaea schausi Godman, [1901]; Biol. centr.-amer., Lep. Rhop. 2: 676, 3: pl. 108, f. 11-12; TL: Mexico, Paso de San Juan, Veracruz
- = ; [NL4A], #2074a; Pelham, 2008, J. Res. Lepid. 40: 372

Chlosyne definita definita; [NL4A], #2074a; Pelham, 2008, J. Res. Lepid. 40: 372; [P&N23]; [iNaturalist] ; [BOA]
Cinclidia albiplaga; Dyar, 1903, Bull. U.S. nat. Mus. 52: 19

C. d. anastasia (Hemming, 1934) Mexico

Melitaea anastasia Hemming, 1934; Stylops 3 (9): 193 (repl. Melitaea beckeri Godman, [1901]); TL: Mexico, Durango
Melitaea beckeri Godman, [1901]; Biol. centr.-amer., Lep. Rhop. 2: 676, 3: pl. 108, f. 9-10 (preocc. Herrich-Schäffer, 1851); TL: Mexico, Durango
- = ; [NL4A], #2074b; Pelham, 2008, J. Res. Lepid. 40: 372

Chlosyne definita anastasia; [NL4A], #2074b; Pelham, 2008, J. Res. Lepid. 40: 372; [P&N23]; [iNaturalist]

Chlosyne kendallorum Opler, 1999

Mexico. See [maps]

Chlosyne kendallorum Opler, 1999; ; TL: Mexico, Nuevo León

Chlosyne kendallorum; [NL4A], #2084; [iNaturalist] ; [BOA]

The Charidryas species-group

Chlosyne palla (Boisduval, 1852)

British Columbia - California. See [maps] Northern Checkerspot

Melitaea palla Boisduval, 1852; Ann. Soc. ent. Fr. (2) 10 (2): 305

Charidryas palla; [NACL], #4492; [NAB], 584
Chlosyne palla; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name); [EBW]; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 381; [MPG] ; [P&N23]; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [BOA] ; [BOA] ; [BOA] ; [BAMONA]
Lemonias palla; Dyar, 1903, Bull. U.S. nat. Mus. 52: 18

C. p. palla California

Melitaea palla ab. wardi Oberthür, 1914; Étud. Lépid. Comp. 9 (2): 79
- = ; Pelham, 2008, J. Res. Lepid. 40: 382
Melitaea palla ab. stygiana Comstock, 1926; Bull. South. Calif. Acad. Sci. 25 (1): 32; TL: Willow Ranch, Modoc Co., California
- = ; Pelham, 2008, J. Res. Lepid. 40: 382

Chlosyne palla palla; Pelham, 2008, J. Res. Lepid. 40: 382; [P&N23]; [iNaturalist] ; [BOA]

C. p. calydon (Holland, 1931) Colorado

Melitaea calydon Holland, 1931; The Butterfly Book (1931 edn): 125, pl. 58, f. 19-20; TL: Turkey Creek Junction, Colorado
Melitaea calydon Mead, 1875; in Wheeler, Reports upon Zool. coll. (8): 760 (Edw., MS)

Charidryas palla calydon; [NACL], #4492d
Chlosyne palla calydon; Pelham, 2008, J. Res. Lepid. 40: 382; [P&N23]; [iNaturalist]
Melitaea calydon; Strecker, 1878, Synon. Cat. Macroplep.: 126

C. p. eremita (Wright, 1905) California

Melitaea eremita Wright, 1905; Butts. West Coast US: 157 ♀, pl. 19, f. 178 b-c; TL: Lake Co., California [San Rafael, Marin Co.]
Melitaea palla ab. hemifusa Gunder, 1930; Ent. News 41: 17; TL: Cazadero, Sonoma Co., California
- = ; Pelham, 2008, J. Res. Lepid. 40: 381

Chlosyne palla eremita; Pelham, 2008, J. Res. Lepid. 40: 381; [P&N23]; [iNaturalist] ; [BOA]

C. p. flavula (Barnes & McDunnough, 1918) Colorado

Melitaea flavula Barnes & McDunnough, 1918; Contr. nat. Hist. Lepid. N. Am. 4 (2): 73, pl. 11, f. 14-15; TL: Glenwood Springs, Colorado

Charidryas palla flavula; [NACL], #4492c
Chlosyne gabbii flavula; [LepIndex]; [CoL]
Chlosyne palla flavula; Pelham, 2008, J. Res. Lepid. 40: 382; [P&N23]; [iNaturalist]

C. p. altasierra Emmel, Emmel & Mattoon, 1998 California

Chlosyne (Charidryas) palla altasierra Emmel, Emmel & Mattoon, 1998; Syst. W. N.Am. Butts. (12): 140; TL: California, El Dorado Co., Fallen Leaf Lake, 6400'
Melitaea (Lemonias) palla r. whitneyi tr.f. vanduzeei Gunder, 1928; Can. Ent. 60 (7): 165; TL: Portola, California
- = ; Pelham, 2008, J. Res. Lepid. 40: 382

Chlosyne palla altasierra; Pelham, 2008, J. Res. Lepid. 40: 382; [P&N23]; [iNaturalist] ; [BOA]

C. p. australomontana Emmel, Emmel & Mattoon, 1998 California

Chlosyne (Charidryas) palla australomontana Emmel, Emmel & Mattoon, 1998; Syst. W. N.Am. Butts. (12): 141; TL: California, Tulare Co., Kennedy Meadows Rd

Chlosyne palla australomontana; Pelham, 2008, J. Res. Lepid. 40: 382; [P&N23]; [iNaturalist] ; [BOA]

C. p. blackmorei Pelham, 2008 SE.British Columbia, E.Washington, NE.Oregon, W.Montana

Chlosyne palla blackmorei Pelham, 2008; J. Res. Lepid. 40: 381; TL: Lytton, British Columbia
Melitaea palla ab. ♀ blackmorei Gunder, 1926; Ent. News 37 (1): 6; TL: Lytton, British Columbia
- = ; Pelham, 2008, J. Res. Lepid. 40: 382

Chlosyne palla blackmorei; [P&N23]; [iNaturalist]

Chlosyne damoetas (Skinner, 1902)

-. See [maps] Rockslide Checkerspot

Melitaea damoetas Skinner, 1902; Ent. News 13 (10): 304; TL: Colorado

Charidryas damoetas; [NACL], #4496; [NAB], 587
Chlosyne damaetas; [LepIndex] ; [CoL]
Chlosyne damaetas[sic, damoetas]; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name)
Chlosyne damoetas; Pelham, 2008, J. Res. Lepid. 40: 382; [MPG] ; [P&N23]; [iNaturalist] ; [BOA]

C. d. damoetas Colorado

Chlosyne damoetas damoetas; Pelham, 2008, J. Res. Lepid. 40: 382; [P&N23]; [iNaturalist] ; [BOA]

C. d. windriver Scott, 1998 Wyoming

Chlosyne whitneyi windriver Scott, 1998; Papilio (n.s.) 11: 3; TL: nr Bears Ears Mtn, 11000', Wind River Mts, Wyoming

Chlosyne damoetas windriver; Pelham, 2008, J. Res. Lepid. 40: 382; [P&N23]; [iNaturalist] ; [BOA]

C. d. altalus Scott, 1998 Alberta, British Columbia

Chlosyne whitneyi altalus Scott, 1998; Papilio (n.s.) 11: 3; TL: Nigel Pass
Melitaea damoeteas ab. damoetella McDunnough, 1927; Can. ent. 59 (7): 160; TL: Mt McLean
- = ; Pelham, 2008, J. Res. Lepid. 40: 383

Chlosyne damoetas altalus; Pelham, 2008, J. Res. Lepid. 40: 383; [P&N23]; [iNaturalist] ; [BOA]

Chlosyne whitneyi (Behr, 1863)

California - Nevada. See [maps]

Melitaea whitneyi Behr, 1863; Proc. Calif. Acad. Sci. 3 (1): 88; TL: California
Melitaea malcolmi Comstock, 1926; Bull. South. Calif. Acad. Sci. 25 (1): 33; TL: nr Mammoth Camp, Mno Co., California
- = ; Pelham, 2008, J. Res. Lepid. 40: 379

Charidryas damoetas malcolmi; [NACL], #4496a
Charidryas palla whitneyi; [NACL], #4492a
Chlosyne malcolmi; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name); [LepIndex] ; [CoL]
Chlosyne palla whitneyi; [LepIndex]; [CoL]
Chlosyne whitneyi; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 379; [MPG] ; [P&N23]; [iNaturalist] ; [BOA] ; [BAMONA]
Lemonias whitneyi; Dyar, 1903, Bull. U.S. nat. Mus. 52: 18

Chlosyne acastus (Edwards, 1874)

Utah, Nevada, Montana. See [maps] Sagebrush Checkerspot

Melitaea acastus Edwards, 1874; Trans. amer. ent. Soc. 5 (1): 16

Charidryas acastus; [NACL], #4493; [NAB], 585
Chlosyne acastus; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name); [BOW]: pl. 13, f. 9; [Opler]; [NL4A], #2071; Pelham, 2008, J. Res. Lepid. 40: 379; [MPG] ; [P&N23]; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [BOA] ; [BOA] ; [BAMONA]
Lemonias acastus; Dyar, 1903, Bull. U.S. nat. Mus. 52: 18

C. a. acastus Montana, Nevada

Chlosyne acastus acastus; Pelham, 2008, J. Res. Lepid. 40: 380; [P&N23]; [iNaturalist] ; [BOA]

C. a. sterope (Edwards, 1870) Oregon, Washington

Melitaea sterope Edwards, 1870; Trans. amer. ent. Soc. 3: 190
Melitaea hewesi Leussler, 1931; Ent. News 42: 12; TL: Tygh, Oregon
- = ; Pelham, 2008, J. Res. Lepid. 40: 380
Melitaea sterope tr.f. hopfingeri Gunder, 1934; Can. Ent. 66 (6): 129; TL: Aster Island, nr Brewster, Okanogan Co., Washington
- = ; Pelham, 2008, J. Res. Lepid. 40: 380

Charidryas palla sterope; [NACL], #4492e
Chlosyne acastus sterope; Pelham, 2008, J. Res. Lepid. 40: 380; [iNaturalist]
Chlosyne palla sterope; [P&N23]
Chlosyne sterope hopfingeri; [LepIndex]; [CoL]
Chlosyne sterope; [LepIndex] ; [CoL]
Lemonias sterope; Dyar, 1903, Bull. U.S. nat. Mus. 52: 18

C. a. neumoegeni (Skinner, 1895) Utah, California Desert Checkerspot

Melitaea neumoegeni Skinner, 1895; Entomol. News 6 (4): 113; TL: Utah
Melitaea neumoegeni tr.f. fridayi Gunder, 1932; Can. Ent. 64 (12): 283; TL: California
- = ; [NL4A], #2071a; Pelham, 2008, J. Res. Lepid. 40: 379
Melitaea neumoegeni tr.f. boharti Gunder, 1933; Can. Ent. 65 (8): 173; TL: Little Rock, Los Angeles Co., California
- = ; Pelham, 2008, J. Res. Lepid. 40: 380

Charidryas neumoegeni; [NACL], #4494; [NAB], 586
Chlosyne acastus neumoegeni; [NL4A], #2071a; Pelham, 2008, J. Res. Lepid. 40: 379; [P&N23]; [LepIndex]; [CoL] ; [iNaturalist] ; [BOA]
Chlosyne neumeogeni[sic]; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name, missp.)
Lemonias neumoegeni; Dyar, 1903, Bull. U.S. nat. Mus. 52: 18

C. a. sabina (Wright, 1905) Arizona

Melitaea sabina Wright, 1905; Butts. West Coast US: 55; TL: Arizona, Catalina Mtns

Charidryas neumoegeni sabina; [NACL], #4494a
Chlosyne acastus sabina; [NL4A], #2071b; Pelham, 2008, J. Res. Lepid. 40: 379; [P&N23]; [iNaturalist] ; [BOA]
Melitaea sabina; [LepIndex] ; [CoL]

C. a. vallismortis (Johnson, 1938) California

Melitaea palla vallis-mortis Johnson, 1938; Bull. S. Calif. Acad. Sci. 37: 18; TL: Tuber Canyon, Panamint Mtns, 8000ft, Death Valley Nat. Monument

Charidryas palla vallismortis; [NACL], #4492b
Chlosyne acastus vallismortis; Pelham, 2008, J. Res. Lepid. 40: 380; [P&N23]; [iNaturalist] ; [BOA]
Chlosyne palla vallismortis; [LepIndex]; [CoL]

C. a. dorothyi Bauer, 1975 Oregon

Chlosyne (Charidryas) acastus dorothyi Bauer, 1975; Butterflies of North America: 157, pl. 40, f. 9

Chlosyne acastus dorothyi; Pelham, 2008, J. Res. Lepid. 40: 380; [P&N23]; [iNaturalist] ; [BOA]

C. a. waucoba Emmel, Emmel & Mattoon, 1998 California

Chlosyne (Charidryas) acastus waucoba Emmel, Emmel & Mattoon, 1998; Syst. W. N.Am. Butts. (12): 142; TL: California, Inyo Co., Inyo Mtns, 8100'
Melitaea acastus ab. ♀ pearlae Gunder, 1926; Ent. News 37 (1): 5; TL: Casa Diablo Hills, Mono Co., California
- = ; Pelham, 2008, J. Res. Lepid. 40: 380

Chlosyne acastus waucoba; Pelham, 2008, J. Res. Lepid. 40: 380; [P&N23]; [iNaturalist] ; [BOA]

C. a. robusta Austin, 1998 Nevada

Chlosyne acastus robusta Austin, 1998; Syst. W. N. Am. Butts. (46): 576; TL: Nevada, Clark Co., Spring Mtns, Kyle Canyon Campground

Chlosyne acastus robusta; Pelham, 2008, J. Res. Lepid. 40: 380; [P&N23]; [iNaturalist] ; [BOA]

C. a. arkanyon Fisher & Scott, 2006 Colorado

Chlosyne strope (acastus) arkanyon Fisher & Scott, 2006; Papilio (n.s.) 12: 37; TL: Poncha Creek, along Hwy 285, S of Poncha Springs, 7800ft, Chaffee Co., Coloarado

Chlosyne acastus arkanyon; Pelham, 2008, J. Res. Lepid. 40: 380; [P&N23]; [iNaturalist] ; [BOA]

Chlosyne gabbii (Behr, 1863)

W.USA. See [maps] Gabb's Checkerspot

Melitaea gabbii Behr, 1863; Proc. Calif. Acad. Sci. 3 (1): 89; TL: California

Charidryas gabbii; [NACL], #4495; [NAB], 587
Chlosyne gabbii; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name); [EBW]; [BOW]: pl. 13, f. 10; [Opler]; [NL4A], #2079; Pelham, 2008, J. Res. Lepid. 40: 380; [MPG] ; [P&N23]; [LepIndex] ; [CoL] ; [iNaturalist] ; [BOA] ; [BAMONA]
Lemonias gabbii; Dyar, 1903, Bull. U.S. nat. Mus. 52: 18

C. g. gabbii California

Melitaea pola Boisduval, 1869; Ann. Soc. ent. Belg. 12: 56; TL: [California]
- = ; [NL4A], #2079a; Pelham, 2008, J. Res. Lepid. 40: 381
Melitaea sonorae Boisduval, 1869; Ann. Soc. ent. Belg. 12: 56; TL: [California]
- = ; [NL4A], #2079a; Pelham, 2008, J. Res. Lepid. 40: 381
Euphydryas gabbii ab. pasadenae Gunder, 1924; Ent. News 35 (5): 155; TL: California
- = ; [NL4A], #2079a; Pelham, 2008, J. Res. Lepid. 40: 381
Melitaea gabbii ab. gunderi Comstock, 1926; Bull. South. Calif. Acad. Sci. 25 (1): 33; TL: California
- = ; [NL4A], #2079a; Pelham, 2008, J. Res. Lepid. 40: 381
Melitaea gabbii ab. newcombi Comstock, 1926; Bull. South. Calif. Acad. Sci. 25 (1): 33; TL: California
- = ; [NL4A], #2079a; Pelham, 2008, J. Res. Lepid. 40: 381

Chlosyne gabbii gabbii; [NL4A], #2079a; Pelham, 2008, J. Res. Lepid. 40: 380; [P&N23]; [iNaturalist] ; [BOA]
Poladryas pola; [LepIndex] ; [CoL]

Misidentified/misapplied:

Melitaea pola (= Poladryas arachne); Bauer, 1975, How Know Butts: 109
?Poladryas pola (= Poladryas arachne?); Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name)

C. g. atrifasciata Hawks & Emmel, 1998 California

Chlosyne (Charidryas) gabbii atrifasciata Hawks & Emmel, 1998; Syst. W. N.Am. Butts. (25): 322; TL: Santa Cruz Island, California

Chlosyne gabbii atrifasciata; Pelham, 2008, J. Res. Lepid. 40: ; [P&N23]; [iNaturalist]

Chlosyne hoffmanni (Behr, 1863)

-. See [maps] Hoffman's Checkerspot Aster Checkerspot

Melitaea hoffmanni Behr, 1863; Proc. Calif. Acad. Sci. 3 (1): 89; TL: California

Charidryas hoffmanni; [NACL], #4497; [NAB], 589
Chlosyne hoffmanni; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name); [BOW]: pl. 13, f. 12; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 378; [MPG] ; [P&N23]; [LepIndex] ; [CoL] ; [iNaturalist] ; [BOA] ; [BAMONA]
Lemonias hoffmanni; Dyar, 1903, Bull. U.S. nat. Mus. 52: 18

C. h. hoffmanni California

Melitaea helcita Boisduval, 1869; Ann. Soc. ent. Belg. 12: 55
- = ; Pelham, 2008, J. Res. Lepid. 40: 378
Melitaea hoffmanni v. abnorma Wright, 1905; Butts. West Coast US: 160, pl. 20, f. 183
- = ; Pelham, 2008, J. Res. Lepid. 40: 379
Melitaea hoffmanni tr.f. hollandae Gunder, 1928; Can. Ent. 60 (7): 166; TL: Summit, California
- = ; Pelham, 2008, J. Res. Lepid. 40: 379

Chlosyne hoffmanni abnorma; [LepIndex]; [CoL]
Chlosyne hoffmanni helcita; [LepIndex]; [CoL]
Chlosyne hoffmanni hoffmanni; Pelham, 2008, J. Res. Lepid. 40: 378; [P&N23]; [BOA]
Chlosyne hoffmanni hollandae; [LepIndex]; [CoL]
Lemonias hoffmanni helcita; Dyar, 1903, Bull. U.S. nat. Mus. 52: 18

C. h. mirabilis (Wright, 1905) California

Melitaea hoffmanni v. mirabilis Wright, 1905; Butts. West Coast US: 160
Melitaea hoffmanni v. mirabilis Wright, 1906; Butts. West Coast (2nd edn.): 160

Chlosyne hoffmanni mirabilis; [LepIndex]; [CoL]

C. h. segregata (Barnes & McDunnough, 1918) Oregon

Melitaea hoffmanni var. segregata Barnes & McDunnough, 1918; Contr. nat. Hist. Lepid. N. Am. 4 (2): 72, pl. 11, f. 8-10; TL: Crater Lake, Oregon, 6500ft
Melitaea bridgei Comstock, 1924; Bull. South. Calif. Acad. Sci. 23 (6): 173; TL: Crater Lake, Oregon
- = ; Pelham, 2008, J. Res. Lepid. 40: 379

Charidryas hoffmanni segregata; [NACL], #4497a
Chlosyne hoffmanni segregata; Pelham, 2008, J. Res. Lepid. 40: 379; [P&N23]; [LepIndex]; [CoL] ; [iNaturalist] ; [BOA]

C. h. manchada Bauer, [1960] Washington

Chlosyne hoffmanni manchada Bauer, [1960]; J. Lep. Soc. 13 (4): 208; TL: 1600ft, Tumwater Canyon, Drury, Chelan Co., Washington

Charidryas hoffmanni manchada; [NACL], #4497b
Chlosyne hoffmanni manchada; Pelham, 2008, J. Res. Lepid. 40: 379; [P&N23]; [LepIndex]; [CoL] ; [iNaturalist] ; [BOA]

The harrisii species-group

Chlosyne harrisii (Scudder, 1863)

NE.USA, Illinois. See [maps] Harris' Checkerspot

Melitaea harrisii Scudder, 1863; Proc. Essex Inst. 3: 167

Charidryas harrisii; [NACL], 61; [NAB], 583
Chlosyne harrisii; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name); [BOW]: pl. 13, f. 11; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 378; [MPG] ; [P&N23]; [LepIndex] ; [CoL] ; [iNaturalist] ; [BOA] ; [BAMONA]
Cinclidia harrisii; Dyar, 1903, Bull. U.S. nat. Mus. 52: 18

C. h. harrisii Maine

Chlosyne harrisii harrisii; Pelham, 2008, J. Res. Lepid. 40: 378; [P&N23]; [iNaturalist] ; [BOA]

C. h. hanhami (Fletcher, 1904) Manitoba, Minnesota Hanham's Crescent

Phyciodes hanhami Fletcher, 1904; Can. Ent. 36 (5): 122

Charidryas harrisii hanhami; [NACL], 61
Chlosyne harrisii hanhami; Pelham, 2008, J. Res. Lepid. 40: 378; [P&N23]; [LepIndex]; [CoL] ; [iNaturalist] ; [BOA]

C. h. liggetti (Avivoff, 1930) Pennsylvania

Melitaea harrisi liggetti Avinoff, 1930; Ann. Carnegie Mus. 19: 161, pl. 6, f. 4-7

Charidryas harrisii liggetti; [NACL], 61
Chlosyne harrisii liggetti; Pelham, 2008, J. Res. Lepid. 40: 378; [P&N23]; [LepIndex]; [CoL] ; [iNaturalist] ; [BOA]

C. h. albimontana (Avinoff, 1930) New Hampshire

Melitaea harrisi albimontana Avinoff, 1930; Ann. Carnegie Mus. 19: 163, pl. 6, f. 8-9; TL: White Mtns

Chlosyne harrisii albimontana; Pelham, 2008, J. Res. Lepid. 40: 378; [P&N23]; [iNaturalist] ; [BOA]

Chlosyne gorgone (Hübner, [1810])

-. See [maps] Gorgone Crescentspot

Dryas gorgone Hübner, [1810]; Samml. exot. Schmett. 1: pl. 41

Charidryas gorgone; [NACL], #4489; [NAB], 580
Chlosyne gorgone; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name); [Opler]; [NL4A], #2081; Pelham, 2008, J. Res. Lepid. 40: 376; [MPG] ; [P&N23]; [LepIndex] ; [CoL] ; [iNaturalist] ; [BOA] ; [BAMONA]

C. g. gorgone South Carolina, NA.Georgia, ...

Chlosyne gorgone gorgone; [NL4A], #2081a; Pelham, 2008, J. Res. Lepid. 40: 376; [iNaturalist] ; [BOA]

C. g. carlota (Reakirt, 1866)

Eresia carlota Reakirt, 1866; Proc. Ent. Soc. Philad. 6 (1): 141; TL: Rocky Mountains, Colorado Territory
Melitaea ismeria Boisduval & Le Conte, [1835]; Hist. Lép. Am. Sept. (11-22): 168, pl. 46, f. 1, 7, 9, 12, 16 [suppr. ICZN 2160]; TL: South Carolina; Georgia
- = Chlosyne gorgone; Calhoun, 2003, J. Lep. Soc. 57 (3): 204-209
- = Chlosyne gorgone gorgone; [NL4A], #2081a
- = ; Pelham, 2008, J. Res. Lepid. 40: 376
Phyciodes ismeria ab. nigra Cary, 1901; Can. Ent. 33: 307; TL: bottom of Monroe Canyon [Sioux Co., Nebraska]
- = ; Pelham, 2008, J. Res. Lepid. 40: 376
nox (Gunder, 1928) (Phyciodes); Can. Ent. 60 (7): 167

Charidryas gorgone carlota; [NACL], #4489a
Charidryas ismeria; Dyar, 1903, Bull. U.S. nat. Mus. 52: 20
Chlosyne gorgone carlota; Iftner, 1983, J. Lep. Soc. 37 (1): 80; Pelham, 2008, J. Res. Lepid. 40: 376; [iNaturalist] ; [BOA]
Chlosyne gorgone nox; [LepIndex]; [CoL]
Melitaea ismeria; [iNaturalist]

C. g. thornei (Gunder, 1934) California

gorgone thornei (Gunder, 1934) (Phyciodes); Can. Ent. 66 (6): 130

Chlosyne gorgone thornei; [LepIndex]; [CoL]

Chlosyne nycteis (Doubleday, [1847])

E.USA. See [maps] Silvery Crescentspot

Melitaea nycteis Doubleday, [1847]; Gen. diurn. Lep. (1): 181, (1): pl. 23, f. 3; TL: USA
?Charidryas harrisii Skinner, 1904

Charidryas nycteis; Dyar, 1903, Bull. U.S. nat. Mus. 52: 20; [NACL], #4490; [NAB], 581
Chlosyne nycteis; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 166 (name); [BOW]: pl. 13, f. 15-16; [Opler]; [NL4A], #2090; Pelham, 2008, J. Res. Lepid. 40: 375; [MPG] ; [P&N23]; [LepIndex] ; [CoL] ; [iNaturalist] ; [BOA] ; [BAMONA]

Misidentified/misapplied:

?Melitaea nycteis; Edwards, 1862, Proc. Acad. nat. Sci. Philad. 13: 161
Phyciodes nycteis (= Phyciodes tharos); Wright, 1906, Butts. West Coast (2nd edn.): 164, pl. 21, f. 197

1025x700 (~167Kb) upperside USA: Alabama, 12.5.2001, Photo © Vitaly Charny	719x906 (~125Kb) underside USA: Alabama, 12.5.2001, Photo © Vitaly Charny	1332x919 (~183Kb) upperside USA: Alabama, 26.5.2001, Photo © Vitaly Charny
742x919 (~86Kb) underside USA: Alabama, 26.5.2001, Photo © Vitaly Charny	1200x782 (~120Kb) upperside USA: Alabama, 12.6.2001, Photo © Vitaly Charny	876x1158 (~90Kb) USA: Alabama, 29.4.2007, Photo © Vitaly Charny
1267x878 (~127Kb) USA: Alabama, 29.4.2007, Photo © Vitaly Charny

C. n. nycteis Massachusetts, Maine, New Jersey, Missouri, Kansas, ...

Melitaea oenone Scudder, 1863; Proc. Essex Inst. 3: 166; TL: Massachusetts; Maine
- = ; [NL4A], #2090a; Pelham, 2008, J. Res. Lepid. 40: 375
Phyciodes nycteis ab. millburni Rummel, 1926; Bull. Brooklyn Ent. Soc. 21: 203; TL: New Jersey
- = ; [NL4A], #2090a
- = (milburni); Pelham, 2008, J. Res. Lepid. 40: 375
Phyciodes nycteis tr.f. lacteus Gunder, 1928; Can. Ent. 60 (7): 166; TL: nr St Louis, Missouri
- = ; [NL4A], #2090a; Pelham, 2008, J. Res. Lepid. 40: 376
Phyciodes nycteis tr.f. greyi Field, 1934; Can. Ent. 66 (11): 256; TL: Kansas
- = ; [NL4A], #2090a; Pelham, 2008, J. Res. Lepid. 40: 376

Chlosyne nycteis greyi; [LepIndex]; [CoL]
Chlosyne nycteis lacteus; [LepIndex]; [CoL]
Chlosyne nycteis nycteis; [NL4A], #2090a; Pelham, 2008, J. Res. Lepid. 40: 375; [P&N23]; [iNaturalist] ; [BOA]

C. n. drusius (Edwards, 1884)

Phyciodes nycteis var. drusius Edwards, 1884; Papilio 4 (3): 57
Phyciodes nycteis drusius tr.f. hewitsoni Field, 1936; J. Ent. Zool. 28 (2): 23; TL: Greaterville, Arizona
- = ; Pelham, 2008, J. Res. Lepid. 40: 376

Charidryas nycteis drusius; Dyar, 1903, Bull. U.S. nat. Mus. 52: 20; [NACL], #4490a
Chlosyne nycteis drusius; Pelham, 2008, J. Res. Lepid. 40: 376; [P&N23]; [LepIndex]; [CoL] ; [iNaturalist] ; [BOA]
Chlosyne nycteis hewitsoni; [LepIndex]; [CoL]

C. n. reversa (Chermock & Chermock, 1940) Manitoba

Phyciodes nycteis reversa Chermock & Chermock, 1940; Can. Ent. 72 (4): 83; TL: Riding Mtns, Manitoba

Charidryas nycteis reversa; [NACL], #4491b
Chlosyne nycteis reversa; Pelham, 2008, J. Res. Lepid. 40: 376; [P&N23]; [CoL] ; [iNaturalist] ; [BOA]

C. n. pastoron Pelham, 2008 North Carolina

Chlosyne nycteis pastoron Pelham, 2008; J. Res. Lepid. 40: 375 (repl. Chlosyne ismeria obsoleta Gatrelle, 2003); TL: North Carolina, Macon Co., Jones Knob 4200
Chlosyne ismeria obsoleta Gatrelle, 2003; (preoc. _Melitaea leanira var. obsoleta Edwards, 1877); TL: North Carolina, Macon Co., Jones Knob 4200

Chlosyne nycteis pastoron; [P&N23]; [BOA]

The Thessalia species-group

Chlosyne leanira (C. & R. Felder, 1860)

Mexico, California. See [maps] Leanira Checkerspot

Melitaea leanira C. & R. Felder, 1860; Wien. ent. Monats. 4 (4): 106 (Boisd. i.l.); TL: California

Chlosyne leanira; [Opler]; [NL4A], #2086; Pelham, 2008, J. Res. Lepid. 40: 374; [MPG] ; [P&N23]; [iNaturalist] ; [BOA] ; [BAMONA]
Phyciodes leanira; [BOW]: pl. 15, f. 1
Thessalia leanira; Dyar, 1903, Bull. U.S. nat. Mus. 52: 19; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name); [NACL], #4505; [NAB], 593; [LepIndex] ; [CoL]

C. l. leanira

Thessalia daviesi Wind, 1947; Pan-Pacific Ent. 23 (4): 171; TL: Strawberry Lake, Tuolumme Co., California, 5500ft
- = ; Pelham, 2008, J. Res. Lepid. 40: 374

Chlosyne leanira leanira; Pelham, 2008, J. Res. Lepid. 40: 374; [P&N23]; [iNaturalist] ; [BOA]

C. l. obsoleta (H. Edwards, 1877) California

Melitaea leanira var. obsoleta H. Edwards, 1877; Proc. Calif. Acad. Sci. 7 (12): 171; TL: nr San Rafael, Marin Co.
Melitaea leanira v. leona Wright, 1905; Butts. West Coast US: 160
- = ; Pelham, 2008, J. Res. Lepid. 40: 374

Chlosyne leanira obsoleta; Pelham, 2008, J. Res. Lepid. 40: 374; [P&N23]; [iNaturalist] ; [BOA]
Melitaea leanira var. obliterata[sic, obsoleta?]; Strecker, 1878, Synon. Cat. Macroplep.: 125
Thessalia leanira obsoleta; Dyar, 1903, Bull. U.S. nat. Mus. 52: 19

C. l. oregonensis (Bauer, 1975) Oregon

Thessalia leanira oregonensis Bauer, 1975; Butterflies of North America: 166, pl. 40, f. 17; TL: Mt Ashland, Loop Road, Jackson Co., Oregon

Chlosyne leanira oregonensis; Pelham, 2008, J. Res. Lepid. 40: 374; [P&N23]; [iNaturalist] ; [BOA]
Thessalia leanira oregonensis; [NACL], #4505a

C. l. wrightii (Edwards, 1886) California

Melitaea wrightii Edwards, 1886; Can. Ent. 18 (4): 64; TL: San Bernardino
Melitaea cerrita Wright, 1905; Butts. West Coast US: 161, pl. 20, f. 189; TL: California
- = ; [NL4A], #2086b; Pelham, 2008, J. Res. Lepid. 40: 375
Melitaea wrightii ab. carlynae Gunder, 1926; Ent. News 37 (1): 3; TL: Mint Canyon
- = ; [NL4A], #2086b; Pelham, 2008, J. Res. Lepid. 40: 375
Melitaea wrightii f. pelona Gunder, 1930; Bull. South. Calif. Acad. Sci. 29 (2): 61; TL: Mint Canyon
- = ; [NL4A], #2086b; Pelham, 2008, J. Res. Lepid. 40: 375

Chlosyne leanira wrightii; [NL4A], #2086b; Pelham, 2008, J. Res. Lepid. 40: 375; [P&N23]; [iNaturalist] ; [BOA]
Thessalia leanira cerrita; [NACL], #4505c
Thessalia leanira wrighti; [NACL], #4505b
Thessalia wrighti; Dyar, 1903, Bull. U.S. nat. Mus. 52: 19; [BOW]: pl. 16, f. 13, 15

C. l. alma (Strecker, [1878]) Utah, Arizona

Melitaea alma Strecker, [1878]; Lep. Rhopal. Het. (15): 135, pl. 15, f. 1
Lemonias alma tr.f. koebelei Gunder, 1927; Ent. News. 38 (5): 136
- = ; Pelham, 2008, J. Res. Lepid. 40: 375

Chlosyne leanira alma; Pelham, 2008, J. Res. Lepid. 40: 375; [P&N23]; [iNaturalist] ; [BOA]
Thessalia alma; Dyar, 1903, Bull. U.S. nat. Mus. 52: 19; [BOW]: pl. 16, f. 18
Thessalia leanira alma; [NACL], #4505d

C. l. elegans (Priestaf & Emmel, 1998) California

Thessalia leanira elegans Priestaf & Emmel, 1998; Syst. W. N.Am. Butts. (26): 327; TL: California, San Luis Obispo Co., Oso Flaco Sand Dunes

Chlosyne leanira elegans; Pelham, 2008, J. Res. Lepid. 40: 374; [P&N23]; [iNaturalist] ; [BOA]

C. l. nebularum (Austin & Smith, 1998) California

Thessalia leanira nebularum Austin & Smith, 1998; Syst. W. N.Am. Butts. (26): 335; TL: California, Santa Cruz Co., Highland Way

Chlosyne leanira nebularum; Pelham, 2008, J. Res. Lepid. 40: 374; [P&N23]; [iNaturalist] ; [BOA]

C. l. basinensis (Austin & Smith, 1998) Nevada

Thessalia leanira basinensis Austin & Smith, 1998; Syst. W. N.Am. Butts. (27): 341; TL: Nevada, Lyon Co., Sweetwater Mtns

Chlosyne leanira basinensis; Pelham, 2008, J. Res. Lepid. 40: 375; [P&N23]; [iNaturalist] ; [BOA]

C. l. flavodorsalis (Austin & Smith, 1998) Utah

Thessalia leanira flavodorsalis Austin & Smith, 1998; Syst. W. N.Am. Butts. (27): 342; TL: Utah, San Juan Co., Comb Ridge

Chlosyne leanira flavodorsalis; Pelham, 2008, J. Res. Lepid. 40: 375; [P&N23]; [iNaturalist] ; [BOA]

C. l. austrima (Austin & Smith, 1998) Mexico (Baja California)

Thessalia leanira austrima Austin & Smith, 1998; ; TL: Mexico, Baja California (Norte)

Chlosyne leanira austrima; [NL4A], #2086a; [iNaturalist] ; [BOA]

Chlosyne cyneas (Godman & Salvin, 1878)

-. See [maps] Cyneas Checkerspot

Phyciodes cyneas Godman & Salvin, 1878; Proc. zool. Soc. Lond. 1878 (1): 269; TL: Mexico, Oaxaca

Chlosyne cyneas; [Opler]; [NL4A], #2073; Pelham, 2008, J. Res. Lepid. 40: 373; [MPG] ; [P&N23]; [iNaturalist] ; [LEPIFORUM] ; [BOA] ; [BAMONA]
Phyciodes cyneas; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 191, 3: pl. 21, f. 10-11; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 677
Thessalia cyneas; Dyar, 1903, Bull. U.S. nat. Mus. 52: 19; [NACL], #4507; [NAB], 595; [BOW]: pl. 16, f. 14

C. c. cyneas Arizona, Mexico

Melitaea (Lemonias) cyneas tr.f. infrequens Gunder, 1928; Can. Ent. 60 (7): 166; TL: Arizona
- = ; [NL4A], #2073a
- = Chlosyne cyneas; Pelham, 2008, J. Res. Lepid. 40: 373

Chlosyne cyneas cyneas; [NL4A], #2073a

C. c. cynisca (Godman & Salvin, 1882) Mexico

Phyciodes cynisca Godman & Salvin, [1882]; Biol. centr.-amer., Lep. Rhop. 1: 191, 3: pl. 21, f. 12; TL: Mexico, Orizaba; Jalapa

Chlosyne cyneas cynisca; [NL4A], #2073b
Chlosyne cynisca; [iNaturalist]
Thessalia cynisca; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name)

Chlosyne theona (Ménétriés, 1855)

-. See [maps] Theona Checkerspot

Melitaea theona Ménétriés, 1855; Cat. lep. Petersb. 2: 86; TL: Nicaragua

Chlosyne theona; [Opler]; [NL4A], #2092; Pelham, 2008, J. Res. Lepid. 40: 373; [MPG] ; [P&N23]; [iNaturalist] ; [LEPIFORUM] ; [BOA] ; [BAMONA]
Phyciodes theona; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 192; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 677
Thessalia theona; Dyar, 1903, Bull. U.S. nat. Mus. 52: 19; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3): 167 (name); [NACL], #4508; [NAB], 596; [BCR], 202; [LepIndex] ; [CoL]

C. t. theona Nicaragua

Chlosyne theona theona; [NL4A], #2092a; [iNaturalist] ; [BOA]

C. t. perlula (C. & R. Felder, 1861) Venezuela

Eresia perlula C. & R. Felder, 1861; Wien. ent. Monats. 5 (4): 104; TL: Venezuela

Chlosyne perlula; [iNaturalist]
Chlosyne theona perlula; [NL4A], #2092j; [P&N23]

C. t. ezra (Hewitson, 1864) Costa Rica, Panama, Honduras

Eresia ezra Hewitson, 1864; Ill. exot. Butts [2] (Eresia III-VI): [66], pl. [34], f. 29
Eresia yorita Reakirt, [1866]; Proc. Ent. Soc. Philad. 5 (2): 224; TL: Honduras
- = ; [NL4A], #2092f

Chlosyne ezra; [iNaturalist]
Chlosyne theona ezra; [NL4A], #2092f
Phyciodes ezra; Godman & Salvin, [1882], Biol. centr.-amer., Lep. Rhop. 1: 192; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 677
Thessalia ezra; [BCR], 202

C. t. thekla (Edwards, 1870) California - Texas, Arizona

Melitaea thekla Edwards, 1870; Trans. amer. ent. Soc. 3: 191; TL: Arizona
tr.f. benjamini (Gunder, 1928) (Melitaea); Can. Ent. 60 (7): 166; TL: Arizona
- = ; [NL4A], #2092k; Pelham, 2008, J. Res. Lepid. 40: 373

Chlosyne thekla; [LepIndex] ; [CoL]
Chlosyne theona thekla; [NL4A], #2092k; Pelham, 2008, J. Res. Lepid. 40: 373; [P&N23]; [iNaturalist] ; [BOA]
Thessalia thekla; Dyar, 1903, Bull. U.S. nat. Mus. 52: 19
Thessalia theona thekla; [NACL], #4508a

C. t. bollii (Edwards, 1877) Texas, Arizona

theona bollii (Edwards, 1877) (Melitaea); Field & Forest 3: 101; TL: Texas

Chlosyne theona bollii; [NL4A], #2092b; Pelham, 2008, J. Res. Lepid. 40: 373; [P&N23]; [iNaturalist]
Thessalia bollii; Dyar, 1903, Bull. U.S. nat. Mus. 52: 19
Thessalia theona bollii; [NACL], #4508b

C. t. hondana (Weymer, 1890) Colombia

Phyciodes hondana Weymer, 1890; Reisen Süd-America: 119, pl. 3, f. 5; TL: Colombia

Chlosyne theona hondana; [NL4A], #2092g; [iNaturalist] ; [BOA]

C. t. brocki (Austin & Smith, 1998) Mexico

Thessalia theona brocki Austin & Smith, 1998; ; TL: Mexico, Sonora

Chlosyne theona brocki; [NL4A], #2092c; [iNaturalist] ; [BOA]

C. t. costaricensis (Austin & Miller, 1998) Costa Rica

Thessalia theona costaricensis Austin & Miller, 1998; ; TL: Costa Rica

Chlosyne theona costaricensis; [NL4A], #2092e; [BOA]

C. t. minimus (Austin & Smith, 1998) Guatemala

Thessalia theona minimus Austin & Smith, 1998; ; TL: Guaetemala

Chlosyne theona minimus; [NL4A], #2092h; [iNaturalist] ; [BOA]

C. t. mullinsi (Austin & Smith, 1998) Mexico

Thessalia theona mullinsi Austin & Smith, 1998; ; TL: Mexico, Oaxaca

Chlosyne theona mullinsi; [NL4A], #2092i; [iNaturalist] ; [BOA]

Chlosyne chinatiensis (Tinkham, 1944)

Texas. See [maps] Chinati Checkerspot

Melitaea chinatiensis Tinkham, 1944; Can. Ent. 76 (1): 12; TL: Texas

Chlosyne chinatiensis; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 26; [MPG] ; [P&N23]; [iNaturalist] ; [BOA]
Chlosyne theona chinatiensis; [NL4A], #2092d; Pelham, 2008, J. Res. Lepid. 40: 373
Thessalia chinatiensis; [NACL], #4509; [NAB], 597

Chlosyne fulvia (Edwards, 1879)

-. See [maps] Fulvia Checkerspot

Melitaea fulvia Edwards, 1879; Can. Ent. 11 (6): 117; TL: Texas

Chlosyne fulvia; [Opler]; [NL4A], #2078; Pelham, 2008, J. Res. Lepid. 40: 374; [MPG] ; [P&N23]; [iNaturalist] ; [LEPIFORUM] ; [BOA] ; [BAMONA]
Thessalia fulvia; [NACL], #4506; [NAB], 594

C. f. fulvia Texas

Melitaea fulvia ab. sinefascia Williams, 1914; Ent. News 25 (6): 265
- = ; Pelham, 2008, J. Res. Lepid. 40: 374

Chlosyne fulvia fulvia; [NL4A], #2078a; Pelham, 2008, J. Res. Lepid. 40: 374

Chlosyne coronado (Smith & Brock, 1988)

Arizona. See [maps]

Thessalia fulvia coronado Smith & Brock, 1988; Bull. Allyn Mus. (118): 6; TL: Arizona, Pima Co., summit of Redington Pass, 20km ENE Tucson, Santa Catalina Mtns

Chlosyne coronado coronado; [P&N23]; [iNaturalist]
Chlosyne coronado; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 26; [MPG] ; [P&N23]; [iNaturalist] ; [BOA]
Chlosyne fulvia coronado; [NL4A], #2078b; Pelham, 2008, J. Res. Lepid. 40: 374

C. c. pariaensis (Smith & Brock, 1988) Utah

Thessalia fulvia pariaensis Smith & Brock, 1988; Bull. Allyn Mus. (118): 10; TL: Utah, Kane Co., Cockscomb Ridge

Chlosyne coronado pariaensis; Zhang, Cong, Shen, Opler & Grishin, 2020, Taxonomic Rep. 8 (7): 26; [P&N23]; [iNaturalist] ; [BOA]
Thessalia fulvia pariaensis; Pelham, 2008, J. Res. Lepid. 40: 374

Synchloe analis Herrich-Schäffer, 1865; (nom. nud.)

analis; [NL4A], 253

Synchloe dromippe Herrich-Schäffer, 1865; (nom. nud.)

dromippe; [NL4A], 254

Synchloe najas Herrich-Schäffer, 1865; (nom. nud.)

najas; [NL4A], 254

Synchloe tegea Boisduval, 1870; (nom. nud.)

tegea; [NL4A], 254

Synchloe tyrinthe Doubleday, [1845]; (nom. nud.); TL: Mexico

tyrinthe; [NL4A], 254

References:

[BAMONA] Opler, Pavulaan, Stanford & Pogue
Butterflies and Moths of North America; Mountain Prairie Information Node
[BCA] electronic
BIOLOGIA CENTRALI-AMERICANA
[BHL] The
Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!
[BOA] Interactive Listing of American Butterflies
Listado Interactivo de la Mariposas Americanas; Butterflies of America
[CoL] Catalogue of Life
Bánki, O., Roskov, Y., Döring, M., Ower, G., Hernández Robles, D. R., Plata Corredor, C. A., Stjernegaard Jeppesen, T., Örn, A., Vandepitte, L., Hobern, D., Schalk, P., DeWalt, R. E., Ma, K., Miller, J., Orrell, T., Aalbu, R., Abbott, J., Adlard, R., Aedo, C., et al. (2024). Catalogue of Life Checklist (Version 2024-03-26); https://doi.org/10.48580/dfz8d
[FLMNH] Florida Museum of Natural History
Bulletin of the Allyn Museum
[HBN] Francis Hemming, 1937
HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[JLS]
Journal of the Lepidopterists' Society; the original J. Lep. Soc. articles
[John Shuey]
e-mail: jshuey at tnc.org; Director of Conservation Science; Indiana Office of The Nature Conservancy; http://www.nature.org
[LAMAS-NEO] Gerardo Lamas
Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)
[LEPIFORUM]
Lepiforum; Bestimmung von Schmetterlingen (Lepidoptera) und ihren Präimaginalstadien
[LepIndex] Global Lepidoptera Index
Beccaloni, G., Scoble, M., Kitching, I., Simonsen, T., Robinson, G., Pitkin, B., Hine, A., Lyal, C., Ollerenshaw, J., Wing, P., & Hobern, D. (2024). Global Lepidoptera Index. In O. Bánki, Y. Roskov, M. Döring, G. Ower, D. R. Hernández Robles, C. A. Plata Corredor, T. Stjernegaard Jeppesen, A. Örn, L. Vandepitte, D. Hobern, P. Schalk, R. E. DeWalt, K. Ma, J. Miller, T. Orrell, R. Aalbu, J. Abbott, R. Adlard, C. Aedo, et al., Catalogue of Life Checklist (1.1.24.106); https://doi.org/10.48580/dg4lg-49xk
[MPG]
North American Moth Photographers Group
[NHM-BMW] Natural History Museum
Butterflies and Moths of the World; Generic Names and their Type-species
[Opler] Opler & Warren, 2003
Butterflies of North America. 2. Scientific Names List for Butterfly Species of North America, north of Mexico.
[P&N23] Pohl & Nanz, 2023
Annotated Taxonomic Checklist of the Lepidoptera of North America, north of Mexico
[Vitaly Charny]
Birmingham, Alabama, USA
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group
[iNaturalist]
iNaturalist
[maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.

Some related literature:

Aaron, [1885]
List of a collection of diurnal Lepidoptera from southern Texas Papilio 4 (9/10) : 172-182
Austin, 1998
New subspecies of Nymphalidae (Lepidoptera) from Nevada and Arizona Syst. W. N. Am. Butts. (46) : 573-586
Avinoff, 1930
Two new subspecies of Melitaea harrisi, with remarks upon related forms Ann. Carnegie Mus. 19 : 161-165, pl. 6
Barnes & McDunnough, 1918
Notes and new species Contr. nat. Hist. Lepid. N. Am. 4 (2): 61-208, pl. 11-25, (3): 211-278, pl. 26-40
Bates, 1864
New species of butterflies from Guatemala and Panama, collected by Osbert Salvin and F. du Cane Godman, Esqs. Ent. mon. Mag. 1 (1): 1-6 (1864), (2): 31-35 (1864), (3): 55-59 (1864), (4): 81-85 (1864), (5): 113-116 (1864), (6): 126-131 (1864), (7): 161-164 (1864), (8): 178-180 (1865), (9): 202-205 (1865)
Bauer, [1960]
A new species of Chlosyne (Nymphalidae) from western Mexico J. Lep. Soc. 13 : 165-169
Bauer, [1960]
A new geographical subspecies of Chlosyne hoffmanni (Nymphalidae) from Washington State J. Lep. Soc. 13 (4) : 207-211
Bauer, [1961]
Descriptions of two new Chlosyne (Nymphalidae) from Mexico, with a discussion of related forms J. Lep. Soc. 14 (2) : 148-154
Behr, 1863
On Californian Lepidoptera - No. III Proc. Calif. Acad. Sci. 3 (1) : 84-93
Boisduval, 1852
Lépidoptères de la Californie Ann. Soc. ent. Fr. (2) 10 (2) : 275-324
Boisduval, 1869
Lépidoptères de la Californie Ann. Soc. ent. Belg. 12 : 5-28,37-94
Boisduval, 1870
Considerations sur des Lépidoptères envoyés du Guatemala à M. de l'Orza Considérations Lépid. Guatemala : 1-100
Boisduval & Le Conte, [1835]
Histoire générale et iconographie des lépidoptères et des chenilles de l'Amérique septentrionale Hist. Lép. Am. Sept. (1): 1-16. pl. 1-3 (1829), (2): 17-24, pl. 4-6 (1829), (3/4): 25-40, pl. 7-12 ([1830]), (5/6): 41-56, pl. 13-18 ([1830]), (7/8): 57-80, pl. 19-24 ([1830]), (9/10): 81-100, pl. 25-30 (1833), (11-22): 101-196, pl. 31-65 ([1835]), (23-26): 197-228, pl. 66-78 ([1837])
Brown & Mielke, 1967
Lepidoptera of the Central Brazil Plateau. I. Preliminary list of Rhopalocera J. Lep. Soc. 21 (2) (2): 77-106, (3): 145-168
Butler, 1870
A revision of the Genera of the Sub-family Pierinae Cistula ent. 1 (3) : 33-58, pl. 1-4
Calhoun, 2003
The history and true identity of Melitaea ismeria (Nymphalidae): a remarkable tale of duplication , misinterpretation, and presumption J. Lep. Soc. 57 (3): 204-219
Cary, 1901
Notes on the butterflies of Sioux county, Nebraska Can. Ent. 33 : 305-311
Chermock & Chermock, 1940
Some new diurnal Lepidoptera from the Riding Mountains and the Sand Ridge, Manitoba Can. Ent. 72 (4) : 81-83
Cockerell, 1894
Synchloe lacinia, aberr. rufescens Can. Ent. 26 (1) : 30
Comstock, 1924
Studies in Pacific Coast Lepidoptera Bull. South. Calif. Acad. Sci. 23 (2): 51-52, (6): 173-176
Comstock, 1926
Studies in Pacific coast Lepidoptera Bull. South. Calif. Acad. Sci. 25 (1) : 29-34, : 48
Doubleday, 1844
List of the Specimens of Lepidopterous insects in The Collection of the British Museum List. lepid. Ins. Brit. Mus. 1 : 1-150
Doubleday, [1847]
The genera of diurnal Lepidoptera, comprising their generic characters, a notice of their habitats and transformations, and a catalogue of the species of each genus; illustrated with 86 plates by W. C. Hewitson Gen. diurn. Lep. (1): 1-18, pl. A, 1-4 (1-2) (1846), (1): 19-132, pl. 4*, 18*, 5-25, 28 (3-14) (1847), (1): 133-200, pl. 1*, 26, 27, 29 (15-23) (1848), (1): pl. 31-44 (15-23) (1848), (1): 201-242, pl. 30, 45-52, 56-58, 60-62, 64 (24-31) (1849), (1): 243-250 [?], (2): 243-326, pl. 53-55, 63, 65, 66 (32-38) (1850), (2): 327-466, pl. 59, 67-77 (39-50) (1851), (2): 467-534, 143, 144, pl. 78-80 (Index) (51-54) (1852)
Drury, 1782
Illustrations of natural history; wherein are exhibited ... Illust. Nat. Hist. Exot. Insects 3 : 1-79 (1837), : pl. 1-50
Dyar, 1903
A List of North American Lepidoptera and Key to the Literature of this Order of Insects Bull. U.S. nat. Mus. 52 : 1-723
Edwards, 1862
Descriptions of certain Species of Diurnal Lepidoptera found within the limits of the United States and of British America. - No. [1]-3 Proc. Acad. nat. Sci. Philad. 13: 160-164 (1861), 14: 54-58, 221-226 (1862)
Edwards, 1870
Descriptions of new North American diurnal Lepidoptera Trans. amer. ent. Soc. 3 : 189-196
Edwards, 1874
Descriptions of new species of diurnal Lepidoptera found in North America Trans. amer. ent. Soc. 5 (1): 13-19, (2): 103-111
Edwards, 1877
Descriptions of new species of diurnal Lepidoptera found in North America Field & Forest 3 (5): 86-89, : 101-105
Edwards, 1879
Description of a new species of Melitaea from Texas Can. Ent. 11 (6) : 117-118
Edwards, 1884
Description. of new species of butterflies mostly from Arizona Papilio 4 (3) : 53-58
Edwards, 1886
Description of new species of butterflies found in the United States Can. Ent. 18 (4) : 61-65
Cockerell in Edwards, 1893; Edwards, 1893
Notes on a polymorphic butterfty, Synchloe lacinia Geyer (in Hüb. Zutr.), with description of its preparatory stages Can. Ent. 25 (11) : 286-291
H. Edwards, 1877
Pacific Coast Lepidoptera. - No. 22. Notes on some Diurnal Lepidoptera, with description of new varieties Proc. Calif. Acad. Sci. 7 (12) : 163-173
Fabricius, 1793
Entomologia Systematica emendata et aucta. Ent. Syst. 3 (1) : 1-487
R. Felder, 1869
Diagnosen neuer von dem K. K. Oberlieutenant H. v. Hedemann in Mexico in den Jahren 1865-1867 gesammelter Lepidopteren. Erste Folge Verh. zool.-bot. Ges. Wien. 19 : 465-480
C. & R. Felder, 1860
Lepidopterologische Fragmente. V-VI Wien. ent. Monats. 4 (4): 97-112, pl. 1-2, (8): 225-251, pl. 3-4
C. & R. Felder, 1861
Lepidoptera nova Columbiae Wien. ent. Monats. 5 (3): 72-87, (4): 97-111
C. & R. Felder, [1867]
Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Behilfen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil. Band 2. Abtheilung 2. Lepidoptera. Rhopalocera Reise Fregatte Novara, Bd 2 (Abth. 2) (1): 1-136, pl. 1-21 (Rhop.) (C. & R. Felder, [1865]), (2): 137-378, pl. 22-47 (Rhop.) (C. & R. Felder, 1865), (3): 379-536, pl. 48-74 (Rhop.) (C. & R. Felder, 1867), (4): 537-548, pl. 75-120, (Inhalts-Verz.) 1-9 (pl. 1-74), (Felder & Rogenhofer, 1874) , (5): pl. 121-140, (erklärung) 1-10 (pl. 75-107), (Inhalts-Verz.) 1-20 (pl. 108-140), (Felder & Rogenhofer, 1875)
Field, 1934
On the naming of "transition forms" in Lepidoptera with notes on certain forms captured near Lawrence, Kansas Can. Ent. 66 (11) : 253-257
Fletcher, 1904
Descriptions of some new species and varieties of Canadian butterflies Can. Ent. 36 (5) : 121-129
Godman & Salvin, 1878
Descriptions of nineteen new species of Diurnal Lepidoptera from Central America Proc. zool. Soc. Lond. 1878 (1) : 264-271
Godman & Salvin, [1882]
Biologia Centrali-Americana. Rhopalocera. Vol. 1. (1879-1886) Biol. centr.-amer., Lep. Rhop. 1: 1-487, 3: pl. 1-24,24a,25-112
Godman & Salvin, [1901]; Godman in Godman & Salvin, [1901]
Biologia Centrali-Americana. Rhopalocera. (1887-1901) Biol. centr.-amer., Lep. Rhop. 2: 1-741 (?782), 3: pl. 1-112
Godman & Salvin, 1894
Descriptions of new Species of Rhopalocera from Mexico and Central America Ann. Mag. nat. Hist. (6) 14 (80) : 95-98
Gunder, 1924
Several New Aberrant Lepidoptera (Rhopalocera) from California Ent. News 35 (5) : 153-158
Gunder, 1926
Several new aberrant Lepidoptera (Rhopalocera) Ent. News 37 (1) : 1-9, pl. 1
Gunder, 1927
New transition forms of "abs" and their classification Ent. News. 38 (5) : 129-138, pl. 2
Gunder, 1928
Additional transitional forms (Lepid., Rhopalocera) Can. Ent. 60 (7) : 162-168
Gunder, 1930
A New Butterfly (Lepid.: Nymphalidae) Ent. News 41 : 17
Gunder, 1930
Butterflies of Los Angeles County, California Bull. South. Calif. Acad. Sci. 29 (2) : 39-95
Gunder, 1932
New Rhopalocera Can. Ent. 64 (12) : 276-284
Gunder, 1933
Additional New Rhopalocera (Lepidoptera) Can. Ent. 65 (8) : 171-173
Gunder, 1934
Various new butterflies (Lepid., Rhopalocera) Can. Ent. 66 (6) : 125-131
Hall, 1917
New butterflies of the family Nymphalidae Entomologist 50 : 161-163, : 171-174
Hall, 1924
Descriptions of two new forms of Chlosyne (Lep. Nymphalidae) Entomologist 57 : 241-242
Hayward, 1931
Lepidopteros Argentinos. Fámilia Nymphalidae Rev. Soc. ent. argent. 4 : 1-199, pl. 1-21
Hayward, 1935
Notas sobre Lepidopteros (Rhop.) Argentinos con description de nuevas especies y formas Rev. Soc. ent. argent. 7: 183-193
Hemming, 1934
Revisional notes on certain species of Rhopalocera (Lepidoptera) Stylops 3 (9) : 193-200
Hewitson, 1864
Illustrations of new species of exotic Butterflies selected chiefly from the collections of W. Wilson Saunders and William C. Hewitson Ill. exot. Butts [2] : (Systematic Index) II, (Ithomia XIV): [1-2], pl. [1] (1855), (Ithomia XV-XIX): [3-12], pl. [2-6] (1856), (Ithomia XX): 13x,14x, [13-14] (1861), pl. [7] (1857), (Ithomia XXI): 15x,16x, [15-16] (1861), pl. [8] (1858), (Ithomia XXII): [17-18], pl. [9] (1859), (Ithomia XXIII): [19-20], pl. [10] (1861), (Ithomia XXIV): [21-22], pl. [11] (1863), (Ithomia XXV): [23-24], pl. [12] (1867), (Ithomia XXVI): [25-26], pl. [13] (1868), (Ithomia XXVII): [27-28], pl. [14] (1869), (Ithomia XXVIII-XXIX): [29-32], pl. [15-16] (1871), (Ithomia XXX-XXXI): [33-36], pl. [17-18] (1872), (Ithomia XXXII-XXXIII): [37-40], pl. [19-20], (1874) , (Ithomia XXXIV-XXXV): [41-44], pl. [21-22] (1876), (Acraea I): [45-46], pl. [23] (1852), (Acraea II): [47-48], pl. [24] (1861), (Acraea III): [49-50], pl. [25] (1865), (Acraea IV-V): [51-52], pl. [26-27] (1867), (Acraea VI): [53-54], pl. [28] (1868), (Acraea VII): [55-56], pl. [29] (1875), (Messaras): [57-58], pl. [30] (1859), (Eresia I): [59-60], pl. [31] (1852), (Eresia II): [61-62], pl. [32] (1857), (Eresia III-VI): [63-70], pl. [33-36] (1864), (Eresia VII): [71-72], pl. [37] (1868), (Eresia VIII-IX): [73-76], pl. [38-39] (1870), (Eresia X): [77-78], pl. [40] (1872), (Junonia): [79-80], pl. [41] (1864), (Cybdelis I-II): [81-84], pl. [42-43] (1852), (Cybdelis III): [85-86], pl. [44] (1857), (Cybdelis IV): [87-88], pl. [45] (1869), (Epiphile I): [89-90], pl. [46] (1857), (Epiphile II & Paromia): [91-92], pl. [47] (1861), (Eubagis I): [93-94], pl. [48] (1852), (Eubagis II): [95-96], pl. [49] (1857), (Eubagis III): [97-98], pl. [50] (1859), (Catagramma I): [99-100], pl. [51] (1851), (Catagramma II): [101-102], pl. [52] (1853), (Catagramma III-V): [103-108], pl. [53-55] (1855), (Catagramma VI): [109-110], pl. [56] (1857), (Catagramma VII-VIII): [111-114], pl. [57-58] (1858), (Catagramma IX and Callithea): [115-116], pl. [59] (1858), (Catagramma X): [117-118], pl. [60] (1864), (Catagramma XI-XII): [119-122], pl. [61-62] (1868), (Catagramma XIII): [123-124], pl. [63] (1876), : iv
Higgins, 1960
A revision of the melitaeine genus Chlosyne and allied species (Lepidoptera: Nymphalidae) Trans. R. ent. Soc. Lond. 112 (14) : 381-475
Higgins, 1981
A revision of Phyciodes Hübner and related genera, with a review of the classification of the Melitaeinae Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 77-243, f. 1-490,124a,125a
[NACL]; Hodges, 1983
Check List of the Lepidoptera of America North of Mexico Check List Lep. Am. N of Mexico
Hübner, [1810]
Sammlung exotischer Schmetterlinge, Vol. 1 (1806 - [1819]) Samml. exot. Schmett. 1 : [1-4] (1807), : [5-12] (1807-1809), : [13-36] (1823-1824), : pl. [1-2], [5-8], [12], [15], [29], [47] (1806), : pl. [9], [11], [17], [21], [25-27], [32], [35], [43-44], [56], [60], [79], [102], [104], [116], [141], [148], [150-151], [155], [159], [200] (1807), : pl. [19], [39], [57], [73], [82], [114], [157], [167], [170], [173], [187] (1807), : pl. [3], [31], [37], [46], [54], [58], [62], [76], [84], [92], [95], [105], [112-113], [117], [137-138], [146], [154], [158], [180], [209] (1808), : pl. [16], [28], [65], [89], [142], [160], [162], [166], [182], [188], [193] (1809), : pl. [4], [22], [42], [53], [61], [88], [90], [97], [100], [107], [110], [118], [120], [140], [143], [149], [161], [171], [178], [184], [196], [202] (1809), : pl. [10], [18], [30], [41], [51], [80], [144], [172], [189] (1810), : pl. [36], [70], [87], [91], [101], [124], [129], [177], [183], [207], [210] (1811), : pl. [20], [38], [45], [55], [85], [98], [103], [108], [136], [139], [152-153], [156], [163], [179], [186], [190], [205-206] (1812), : pl. [13-14], [40], [48], [50], [67], [74], [86], [119], [121], [130], [135], [175-176], [197-199], [204], [212] (1813), : pl. [72], [75], [77-78], [83], [93], [109], [111], [115], [128], [131], [165], [174], [194] (1813), : pl. [24], [49], [52], [63-64], [66], [68-69], [71], [81], [123], [145], [181], [192] (1814), : pl. [126], [132-134], [168] (1815), : pl. [23], [96], [122], [125], [169], [195] (1816), : pl. [33], [99], [127], [187], [185], [213] (1817), : pl. [59], [106], [147], [164], [208], [211] (1818), : pl. [34], [94], [191], [201] (1819), : (Index Systematicus) [1-4] (1819 - April 1823)
Geyer, [1833]
Sammlung exotischer Schmetterlinge, Vol. 3 ([1827] - [1838]) in Hübner, Samml. exot. Schmett. 3 : pl. [9], [11], [21], [26-27], [32], [43], [45] (1827), : pl. [7], [13], [35], [42] (1828), : pl. [20], [22], [37-38] (1829), : pl. [17], [24], [41], [49] (1830), : pl. [30] (1831), : pl. [25], [28], [31], [44], [50-51] (1832), : pl. [5-6], [14-16], [29], [48], [52], [54] (1833), : pl. [1], [12], [18] (1834), : pl. [2-4], [10], [19], [23], [39], [46] (1835), : pl. [40] (1836-1837), : pl. [8], [33-34], [36], [47], [53] (1838)
Geyer, 1837
Zuträge zur Sammlung exotischer Schmettlinge, Vol. 5 [1833]-1837 in Hübner, Zuträge. Samml. exot. Schmett. 5 : [1-3], 4, [5], 6-52 (1837), : pl. [138-151], f. 801-880 (1833), : pl. [152-172], f. 881-1000 (1834 - 1836)
Iftner, 1983
A new food plant record for Chlosyne gorgone carlota (Reakirt) (Nymphalidae) J. Lep. Soc. 37 (1) : 80-81
Johnson, 1938
A new subspecies of Melitaea palla Boisduval 1852 (Lepidoptera, Nymphalidae) Bull. S. Calif. Acad. Sci. 37 : 18-20
Kirby, 1871
A synonymic catalogue of the diurnal Lepidoptera Syn. Cat. diurn. Lep.: vii + 690pp : 1-690
[NL4A]; Lamas, 2004
Checklist: Part 4A, Hesperioidea - Papilionoidea Atlas of Neotropical Lepidoptera
Latreille, [1809]
Insectes de l'Amérique équinoxiale, recueillis pendant de voyage de MM. de Humboldt et Bonpland. In Humbolt & Bonpland, aux régions équinoxiales du Nouveau Continent. 2. Recueil d'observations de zoologie et d'anatomie comparée; faites dans l'Océan Atlantique, dans l'intérieur du Nouveau Continent et dans la Mer du Sund, pendant les années 1799, 1800, 1801, 1802 et 1803 in Humboldt & Bonpland, Voy. rég. équinoxiales..., 1 (4): 197-283, pl. 15-18 (1809), (5/6): 344-397, pl. 22-25 (1809), (7): 127-252, pl. 15-18,22-25 (1811)
Leussler, 1931
A New Melitaea from Oregon (Lep.: Nymphalidae) Ent. News 42 : 12-13
[BOW]; Lewis, 1974
Butterflies of the World
de la Maza & de la Maza, 2019
Identificación de la población nominal de Chlosyne melitaeoides (C. y R. Felder, [1867]) y descripción de una nueva subespecie de Puebla y Oaxaca, México (Nymphalidae: Melitaeini) Rev. Soc. Mex. Lep. (ns) 6 (2) : 2-15
McDunnough, 1927
The Lepidoptera of the Seton Lake Region, British Columbia Can. ent. 59 (7): 152-162, (8): 193-199, (9): 207-214, (10): 239-246, (11): 266-277
Mead, 1875
Report upon the collections of diurnal Lepidoptera made in portions of Colorado, Utah, New Mexico, and Arizona, during the years 1871, 1872, 1873, and 1874, with notes upon all species known to inhabit Colorado in Wheeler, Reports upon Zool. coll. (8) : 739-794
Ménétriés, 1855
Enumeratio corporum animalium Musei Imperialis Academiae Scientiarum Petropilitanae. Classis Insectorum, Ordo Lepidopterorum Cat. lep. Petersb. 1: 1-66, pl. 1-6, (supplement) 67-112 (1855), 2: 67-97,99-144, pl. 7-14 (1857), 3: 145-161, pl. 15-18 (1863)
Miller & Rotger, 1979
Two new Chlosyne (Nymphalidae) from Mexico Bull. Allyn Mus. 54 : 1-4
Pelham, 2008
A catalogue of the butterflies of the United States and Canada J. Res. Lepid. 40 : 1-652
[NAB]; Pyle, 1981
Field Guide to North American Butterflies, National Audubon Society
[EcuL]; Racheli & Racheli, 2001
An annotated list of Ecuadorian Butterflies (Lepidoptera: Papilionidae, Pieridae, Nymphalidae) Frag. ent. 33 (2) : 213-380
Reakirt, [1866]
Descriptions of some new species of Eresia Proc. Ent. Soc. Philad. 5 (2) : 224-227
Reakirt, 1866
Coloradian butteflies Proc. Ent. Soc. Philad. 6 (1) : 122-151
Reakirt, 1866
Descriptions of some new species of Diurnal Lepidoptera Proc. Acad. nat. Sci. Philad. 18 (3) : 238-249
Reakirt, [1867]
Descriptions of some new species of Diurnal Lepidoptera. Series II Proc. Acad. nat. Sci. Philad. 18 (4): 331-336 (1867), (5): 337-342 (1867)
Rummel, 1926
Notes on aberrations of New Jersey butterflies Bull. Brooklyn Ent. Soc. 21 : 203
Röber, 1914
4. Genus Phyciodes Hbn. - 7. Gattung: Gnatotriche Fldr. in Seitz, Gross-Schmetterl. Erde 5 : 434-440 (10 Dec 1913), : 441-448 (15 Dec 1913), : 449-454 ([before 28 Feb] 1914), : pl. 88-92 (1914)
Scott, 1986
The Butterflies of North America Butts. N. America : 1-583
Scott, 1998
New western North American butterflies Papilio (n.s.) 11 : 1-12
Scudder, 1863
A List of the Butterflies of New England Proc. Essex Inst. 3 : 161-179
Scudder, 1872
A systematic Revision of some of the American Butterflies; with brief notes on those known to occur in Essex County, Mass. 4th Ann. Rep. Peabody Acad. Sci. (1871) : 24-83,errata
Scudder, 1875
Synonymic list of the butterflies of North Amercia, north of Mexico. (1) Nymphales Bull. Buffalo Soc. nat. Sci. 2 : 233-269
Skinner, 1895
Notes on Rhopalocera, with descriptions of new species Entomol. News 6 (4) : 112-114
Skinner, 1902
A New Species of Melitaea Ent. News 13 (10) : 304-305
[EBW]; Smart, 1976
The Illustrated Encyclopedia of the Butterfly World Illust. Encyp. Butt. World: -
Smith & Brock, 1988
A review of the Thessalia leanira complex in the southwestern United States (Nymphalidae: Melitaeinae), with a description of two new subspecies of Thessalia fulvia Bull. Allyn Mus. (118) : 1-21
Staudinger, 1876
Neue Lepidopteren des südamerikanischen Faunengebietes Verh. zool.-bot. Ges. Wien, 25 (1) : 89-124
Staudinger, 1885
Exotische Tagfalter in sysmatischer Reihenfolge mit Berücksichtigung neuer Arten in Staudinger & Schatz, Exot. Schmett. 1 (1): 3-6, pl. 1-5 (1884), (2): 7-10, pl. 6-10 (1884), (3): 11-14, pl. 11-15 (1884), (4): 15-22, pl. 16-20 (1884), (5): 23-30, pl. 21-25 (1884), (6): 31-38, pl. 26-30 (1884), (7): 39-46, pl. 31-35 (1885), (8): 47-58, pl. 30-40 (1885), (9): 59-82, pl. 41-50 (1885), (10): 83-102, pl. 51-60 (1885), (11): 103-122, pl. 61-65 (1886), (12): 123-138, pl. 66-70 (1886), (13): 139-158, pl. 71-75 (1886), (14): 159-174, pl. 76-80 (1886), (15): 175-194, pl. 81-85 (1887), (16): 195-214, pl. 86-90 (1887), (17): 215-234, pl. 91-95 (1887), (18): 235-270, pl. 96-98 (1888), (19): 1-2,271-333, pl. 99-100 (1888)
Strecker, [1878]
Lepidoptera, Rhopaloceres and Heteroceres, indigenous and exotic, with descriptions and colored illustrations Lep. Rhopal. Het. (1): 1-7, pl. 1 (1872), (2): 9-15, pl. 2 (1873), (3): 17-24, pl. 3 (1873), (4): 25-32, pl. 4 (1873), (5): 33-43, pl. 5 (1873), (6): 45-50, pl. 6 (1873), (7): 51-60, pl. 7 (1873), (8): 61-70, pl. 8 (1874), (9): 71-80, pl. 9 (1874), (10): 81-94, pl. 10 (1874), (11): 95-100, pl. 11 (1874), (12): 101-108, pl. 12 (1875), (13): 109-123, pl. 13 (1876), (14): 125-134, pl. 14 (1878), (15): 135-143, pl. 15 (1878)
Strecker, 1878
Butterflies and Moths of North America. Complete Synonymical Catalogue of Macrolepidoptera ... Synon. Cat. Macroplep.: - : 1-281
Tinkham, 1944
Faunistic notes on the diurnal Lepidoptera of the Big Bend region of Trans-Pecos, Texas, with the description of a new Melitaea Can. Ent. 76 (1) : 11-18
[BCR]; DeVries, 1987
Papilionidae, Pieridae, Nymphalidae The Butterflies of Costa Rica 1
Weymer in Weymer & Maassen, 1890
Lepidoptera gesammelt auf einer Reise durch Colombia, Ecuador, Peru, Brasilien, Argentinien und Bolivien in den Jahren 1868-1877 von Alphons Stübel Reisen Süd-America : 1-182, pl. 1-9
Williams, 1914
One Hundred Butterflies from the James Mountains New Mexico (Lepid.), With notes and Description of a New Species Ent. News 25 (6) : 263-268
Wind, 1947
A new subspecies of Melitaea (Lepidoptera) Pan-Pacific Ent. 23 (4) : 171
Wright, 1905
The Butterflies of the West Coast of the United States Butts. West Coast US : 3-257
Wright, 1906
The Butterflies of the west coast of the United States Butts. West Coast (2nd edn.) : 1-257
Zhang, Cong, Shen, Opler & Grishin, 2020
Genomic evidence suggests further changes of butterfly names Taxonomic Rep. 8 (7) : 1-41
Zhang, Cong, Shen, Opler & Grishin, 2021
Genomics-guided refinement of butterfly taxonomy Taxonomic Rep. 9 (3) : 1-55

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