Erynnis

Erynnis Schrank, 1801 Duskywings

Erynnis Schrank, 1801; Fauna Boica 2 (1): 152, 157; TS: Papilio tages Linnaeus
Thymele Fabricius, 1807; Mag. f. Insektenk. 6: 287; TS: Papilio tages Linnaeus
Thymale Oken, 1815; Lehrbuch Naturgesch. 3 (Zool.) (1): 757-758 (unavail.); TS: Papilio tages Linnaeus
Astycus Hübner, 1822; Syst.-alph. Verz.: 1, 3, 5, 6, 8-10; TS: Papilio tages Linnaeus
Thanaos Boisduval, [1834]; Icon. hist. Lépid. Europe 1 (23-24): 240; TS: Papilio tages Linnaeus
Thanatos Dunning & Pickard, 1858; Accent. List Brit. Lep.: 6 (unjust. emend.)
Hallia Tutt, 1906; Nat. Hist. Br. Lepid. 8: 261 (preocc. Hallia Edwards & Haime, 1850); TS: Thanaos marloyi Boisduval
Erynnides Burns, 1964

Erynnis (Erynnides); [P&N23]
Erynnis (Erynnis); [P&N23]
Erynnis; [P&N23]; [LepIndex]; [CoL] ; [iNaturalist] ; [NHM-BMW]
Erynnis (Pyrginae); Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 8; [RFEL]
Erynnis (Pyrgini); [NL4A], 47
Thanaos; Godman & Salvin, [1899], Biol. centr.-amer., Lep. Rhop. 2: 455; [NHM-BMW]
Thanatos; [NHM-BMW] ; [NHM-BMW]
Thymele; [NHM-BMW] ; [NHM-BMW]

900x617 (~75Kb) USA: Near Crystal Mtn, (46°57'51N 121°28'55W), Pierce Co., WA, 27.7.1999, Photo © Markku Savela

700x652 (~66Kb) USA: Near Crystal Mtn, (46°57'51N 121°28'55W), Pierce Co., WA, 27.7.1999, Photo © Markku Savela

identification uncertain 500x378 (~51Kb) Venezuela, Parai Tepui near Mount Roraima in the Gran Sabana, July 2002, Photo © Simon Coombes

The exact identification of these species is still unknown, but tentatively assumed to belong into this group.

[NL4A] introduced subgeneric division, but I don't know how to place the species outside neotropics.

Erynnis montanus (Bremer, 1861) S.China - Amur, Japan. See [maps]
- Hesperia montanus Bremer, 1861; Bull. Acad. Imp. Sci. St. Petersb. 3 Mélanges biol. St.-Pétersb. 3 (5-6): 556; TL: Bureinsky Mts., Amur region
- Thanaos rusticanus Butler, 1866; J. Linn. Soc. Zool. Lond. 9 (34): 58; TL: Japan, Hakodate
- Erynnis montanus; [BOW]: pl. 208, f. 9
- Erynnis montanus; [BRU], 106; [MRS], 705; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 8; [RFEL]; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [SZMN]
- Erynnis mountanus[sic, montanus]; Kudrna, 1974, Atalanta 5: 116
Larva on Quercus mongolica [BRU]
- E. m. nigrescens (Leech)
  - Erynnis montanus nigrescens; [MRS], 705; [LepIndex]; [CoL]
- E. m. monta Evans
  - Erynnis montanus monta; [MRS], 705; [LepIndex]; [CoL]
- E. m. neomontanus Murayama & Yoshisaka, 1959 Formosa
  - Erynnis montanus neomontanus Murayama & Yoshisaka, 1959; Tyô to Ga 10 (1): 14; TL: Poli?, Formosa

Erynnis popoviana (Nordmann, 1851)

Mongolia, N.China, E.China, C.China, Ussuri. See [maps]

Hesperia popoviana Nordmann, 1851; Bull. Soc. imp. Nat. Moscou 24: 443, pl. 12, f. 3-4; TL: Kyakhta, Transbaikalia
Nisoniades tages var. sinina Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4): 461

Erynnis popoviana; [BRU], 107; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 8; [RFEL]; [iNaturalist]
Erynnis tages popoviana; [MRS], 705; [LepIndex]; [CoL]

800x600 (~121Kb) upperside Wet ground at the pool in the pine forest called Tsasucheiskii Bor at the village Nizhnii Tsasuchei, Onon District, Chita Province, SE Transbaikalia, Siberia, Russia. 22nd June 1995, Photo © Oleg Kosterin

Erynnis pathan Evans, 1949 Ghissar - Chitral, N.Baluchistan. See [maps]
- Erynnis pathan Evans, 1949; Cat. Hesp. Europe Asia Australia Brit. Mus.: 168; TL: Chitral, Utzun Valley
- Erynnis marloyi pathan; [LepIndex]; [CoL]
- Erynnis pathan; [BRU], 107; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 8
Erynnis pelias (Leech, 1891) W.China. See [maps]
- Nisoniades pelias Leech, 1891; Entomologist 24 (Suppl.): 60; TL: Ta-chein-lu; Wa-ssu-kou
- Nisoniades erebus Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4): 461
- Erynnis pelias erebus; [LepIndex]; [CoL]
- Erynnis pelias; Huang, 2001; [LepIndex] ; [CoL] ; [iNaturalist]

Subgenus Erynnis Schrank, 1801

Erynnis Schrank, 1801; Fauna Boica 2 (1): 152, 157; TS: Papilio tages Linnaeus
Thymele Fabricius, 1807; Mag. f. Insektenk. 6: 287; TS: Papilio tages Linnaeus
Thymale Oken, 1815; Lehrbuch Naturgesch. 3 (Zool.) (1): 757-758 (unavail.); TS: Papilio tages Linnaeus
Astycus Hübner, 1822; Syst.-alph. Verz.: 1, 3, 5, 6, 8-10; TS: Papilio tages Linnaeus
Thanaos Boisduval, [1834]; Icon. hist. Lépid. Europe 1 (23-24): 240; TS: Papilio tages Linnaeus
Thanatos Dunning & Pickard, 1858; Accent. List Brit. Lep.: 6 (unjust. emend.)
Hallia Tutt, 1906; Nat. Hist. Br. Lepid. 8: 261 (preocc. Hallia Edwards & Haime, 1850); TS: Thanaos marloyi Boisduval

Astyeus; Evans, 1953, (missp.)
Erinnys; Ochsenheimer, 1816, (missp.); [NHM-BMW]
Erymnis; Rafinesque, 1815, (missp.); [NHM-BMW]
Erynnis (Erynnis); [NL4A], 47
Hallia; [NHM-BMW]
Prynnis; Miller, 1992, (missp.); [NHM-BMW]
Tanaus; Blanchard, 1840, (missp.); [NHM-BMW]
Thanaes; Simms, 1920, (missp.); [NHM-BMW]
Thanaus; Kirby, 1897, in Allen, Nat. Libr., Lepid. 3: 12 (missp.); [NHM-BMW]
Thannaos; Lang, 1884, (missp.); [NHM-BMW]
Thimele; Hayward, 1933, (missp.); [NHM-BMW]
Thymale; [NHM-BMW] ; [NHM-BMW]
Thymate; Hayward, 1973, (missp.); [NHM-BMW]
Thymela; Raymundo da Silva, 1907, (missp.); [NHM-BMW]
Thymeles; Nekrutenko, 2003, (missp.); [NHM-BMW]

Erynnis tages (Linnaeus, 1758)

CEU, SEU, tr, Russia, Siberia, C.Asia, China, Amur. See [maps] La Grisette Cervantes Skogssmygare Dunkler Dickopffalter Powszelatek brunatek Dingy Skipper

Papilio tages Linnaeus, 1758; Syst. Nat. (Edn 10) 1: 485
morio Scopoli, 1763
geryon Rottenburg, [1775]

Erynnis tages baynesi; [LepIndex]; [CoL]
Erynnis tages cervantes; [LepIndex]; [CoL]
Erynnis tages; [H&R], 331; [BOW]: pl. 11, f. 15; [BRU], 106; [MRS], 705; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 8; Yakovlev, 2012, Nota lepid. 35 (1): 60; [RFEL]; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [SVF]

500x319 (~36Kb) upperside Haldon woods, Devon, UK 21/05/95, Photo © Simon Coombes

800x600 (~114Kb) upperside Open pine forest on a hill just NE of the Chita City, Transbaikalia, Siberia, Russia. 17th June 1995, Photo © Oleg Kosterin

Larva on Lotus corniculatus, Eryngium , Coronilla [H&R], Medicago , Hippocrepis [BRU]

E. t. unicolor Freyer, 1852 Transcaucasus
- Erynnis tages unicolor; [LepIndex]; [CoL]
E. t. elbursina (Bytinski-Salz & Brandt, 1937) Iran
- Thanaos elbursina Bytinski-Salz & Brandt, 1937; Ent. Rec. J. Var. (Suppl.) 49: 3
- Erynnis tages elbursina; [LepIndex]; [CoL]

Erynnis marloyi (Boisduval, [1834])

Balkans, Asia Minor, Syria, Iran, Chitral. See [maps] Inky Skipper

Thanaos marloyi Boisduval, [1834]; Icon. hist. Lépid. Europe 1 (23-24): 241, pl. 47, f. 6-7; TL: Peloponnesos, Greece
sericea Freyer, [1838]
rustan Kollar, [1849]

Erynnis marloyi max; [LepIndex]; [CoL]
Erynnis marloyi; [H&R], 331; [BOW]: pl. 11, f. 14; [BRU], 107; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 8; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [MBE]

1000x1212 (~415Kb) Iran, Ostan-e Markazi, Shakhrestan-e Shazand, Astaneh Town env., Seidan chashmah, 2289 m a.s.l., 20th May 2017, Photo © Oleg Kosterin

1000x1121 (~410Kb) Iran, Ostan-e Markazi, Shakhrestan-e Shazand, Astaneh Town env., Seidan chashmah, 2289 m a.s.l., 20th May 2017, Photo © Oleg Kosterin

Erynnis brizo (Boisduval & Le Conte, [1834]) E.USA, Colorado, Arizona. See [maps] Sleepy Duskywing
- Thanaos brizo Boisduval & Le Conte, [1834]; Hist. Lép. Am. Sept. (23-26): pl. 66; TL: N.Amer.
- Erynnis (Erynnis) brizo; [NL4A], #751; [P&N23]
- Erynnis brizo; [NACL], #3946; [NAB], 748; [Opler]; [MPG] ; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [BOA] ; [BAMONA]
- Thanaos brizo; Dyar, 1903, Bull. U.S. nat. Mus. 52: 59
Larva on Quercus , less Castanea dentata [NAB]
- E. b. brizo Mexico, ...
  - Erynnis (Erynnis) brizo brizo; [NL4A], #751a
  - Erynnis briso; Okano, 1981, (missp.)
  - Erynnis brizo brizo; [iNaturalist]
- E. b. somnus (Lintner, 1881) Florida?
  - Nisoniades[?] somnus Lintner, 1881; Papilio 1 (5): 73
  - Erynnis (Erynnis) somnus; [P&N23]
  - Erynnis brizo somnus; [NACL], #3946a; [iNaturalist]
  - Erynnis somnus; [MPG]
  - Thanaos somnus; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
- E. b. lacustra (Wright, 1905) Mexico, California
  - brizo lacustra (Wright, 1905) (Nisoniades); ; TL: California
  - Erynnis (Erynnis) brizo lacustra; [NL4A], #751c
  - Erynnis (Erynnis) lacustra; [P&N23]
  - Erynnis brizo lacustra; [NACL], #3946c; [iNaturalist]
  - Erynnis lacustra; [MPG]
  - Erynnis pacuvius lacustra Wright, 1904;
- E. b. burgessi (Skinner, 1914) Mexico, Arizona
  - brizo burgessi (Skinner, 1914) (Thanaos); ; TL: Arizona
  - Erynnis (Erynnis) brizo burgessi; [NL4A], #751b
  - Erynnis (Erynnis) burgessi; [P&N23]
  - Erynnis brizo burgessi; [NACL], #3946b; [iNaturalist]
  - Erynnis burgessi; [MPG]
  - Erynnis burgessii; Klots, 1951, (missp.)
- E. b. muelleri (Draudt, 1923) Mexico
  - brizo muelleri (Draudt, 1923) (Thanaos); ; TL: Mexico, Puebla
  - Erynnis (Erynnis) brizo muelleri; [NL4A], #751d
  - Erynnis brizo muelleri; [iNaturalist]
  - Thanaos mulleri; Hoffmann, 1932, (missp.)
Erynnis icelus (Scudder & Burgess, 1870) New York, ..?.., California. See [maps] Aspen Dusky Wing Dreamy Duskywing
- Nisoniades icelus Scudder & Burgess, 1870; ; TL: "New England"
- Nisoniades icelus Lintner, 1872; 23 Rept. N.Y. Sta. Cab. Nat. Hist.: 162 (preocc.); TL: New York
- Nisoniades bautista Plötz, 1884; ; TL: California
  - = ; [NL4A], #752
- Nisoniades hamamaelidis Scudder, 1889; (nom. nud.); TL: USA
  - = ; [NL4A], #752
- Erynnis (Erynnis) icelus; [NL4A], #752; [P&N23]
- Erynnis hamamaelides; Bridges, 1983, (missp.)
- Erynnis icelus; [NACL], #3945; [NAB], 748; [BOW]: pl. 21, f. 36; [BOC], 51; [Opler]; [MPG] ; [LepIndex] ; [CoL] ; [iNaturalist] ; [BOA] ; [BAMONA]
- ?Thanaos icelus Lintner, 1872; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
Larva on Salix , Populus , Robinia , Betula [NAB]
Erynnis mercurius (Dyar, 1926) Mexico. See [maps]
- Thanaos mercurius Dyar, 1926; ; TL: Mexico, Distrito Federal
- Erynnis (Erynnis) mercurius; [NL4A], #753
- Erynnis mercurius; [iNaturalist] ; [BOA]

Subgenus Erynnides Burns, 1964

Erynnides; [NHM-BMW]
Erynnides (Erynnis); [NL4A], 47

Erynnis afranius (Lintner, 1878) Mexico - Manitoba. See [maps] Bald Duskywing Afranius Duskywing
- Nisoniades afranius Lintner, 1878; 30 Rept. N.Y. Mus. Nat. Hist.: 175; TL: "Colorado"
- Erynnis (Erynnides) afranius; [NL4A], #754; [P&N23]
- Erynnis afranius; [NACL], #3960; [NAB], 759 (note); [BOC], 54; [Opler]; [MPG] ; [LepIndex] ; [CoL] ; [iNaturalist] ; [BAMONA]
- Thanaos afranius; Godman & Salvin, [1899], Biol. centr.-amer., Lep. Rhop. 2: 459, 3: pl. 91, f. 24-26; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
Larva on Lupinus , Lotus [NAB], Thermopsis rhombifolia [BOC]
Erynnis funeralis (Scudder & Burgess, 1870) California, S.Nevada, S.Utah, Colorado, W.Kansas - Texas - Argentina, Chile, Colombia. See [maps] Funereal Duskywing
- Nisoniades funeralis Scudder & Burgess, 1870; Proc. Bost. Soc. Nat. Hist. 13: 293; TL: Texas
- Helias clericalis Burmeister, 1875; ; TL: Argentina
  - = ; [NL4A], #755
- Nisoniades australis Mabille, 1883; Bull. Soc. Ent. Belg. 1883: liv [54]; TL: Colombia
  - = ; [NL4A], #755
- Erynnis (Erynnides) funeralis; [NL4A], #755; [P&N23]
- Erynnis funeralis; [NACL], #3957; [NAB], 736; [BOW]: pl. 21, f. 34; [Opler]; Benyamini, Ugarte, Shapiro, Mielke, Pyrcz & Bálint, 2014, Bol. Mus. Nac. Hist. Nat. Chile 63: 21 (list); [MPG] ; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [BAMONA]
- Erynnis zarucco funeralis;
- Phlebodes clericalis; Burmeister, 1878
- Thanaos funeralis; Godman & Salvin, [1899], Biol. centr.-amer., Lep. Rhop. 2: 456, 3: pl. 91, f. 10-14; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
Larva on Fabaceae [NAB]
Erynnis horatius (Scudder & Burgess, 1870) E.USA, Florida, Mexico. See [maps] Horace's Duskywing
- Nisoniades horatius Scudder & Burgess, 1870; Proc. Bost. Soc. Nat. Hist. 13: 301; TL: Massachusetts/Texas
- Nisoniades virgilius Scudder & Burgess, 1870; ; TL: USA
  - = ; [NL4A], #756
- Nisoniades petronius Mead, 1875; (nom. nud.)
- Nisoniades petronius Lintner, 1881; Papilio 1 (5): 70; TL: Florida
  - = ; [NL4A], #756
- Erynnis (Erynnides) horatius; [NL4A], #756; [P&N23]
- Erynnis horatius; [NACL], #3952; [NAB], 752; [BOW]: pl. 21, f. 35; [Opler]; [MPG] ; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [BAMONA]
- Thanaos horatius; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
- Thanaos horatuis[sic, horatius]; Draudt, 1923
- Thanaos petronius; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
Larva on Quercus muehlenbergii, Q. gambelii, Q. texana [NAB]
Erynnis juvenalis (Fabricius, 1793) E.USA - Rocky Mts.. See [maps] Juvenal's Duskywing
- Hesperia juvenalis Fabricius, 1793; Ent. Syst. 3 (1): 339; TL: America
- Erynnis (Erynnides) juvenalis juvenalis; [P&N23]
- Erynnis (Erynnides) juvenalis; [NL4A], #757; [P&N23]
- Erynnis juvenalis; [NACL], #3947; [NAB], 749; [BOW]: pl. 21, f. 35 (text); [Opler]; [MPG] ; [LepIndex] ; [CoL] ; [iNaturalist] ; [BAMONA]
- Thanaos juvenalis; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
Larva on Quercus velutina, Q. rubra, Q. alba [NAB]
- E. j. juvenalis Florida, ...?
  - Nisoniades juvenis Hübner, [1819]; (repl. name); TL: America
    
    = Thanaos juvenalis; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
    = ; [NL4A], #757a
  - Nisoniades costalis Westwood, 1852
    
    = ; [NL4A], #757a
  - Nisoniades ennius Scudder & Burgess, 1870; ; TL: USA
    
    = ; [NL4A], #757a
  - Nisoniades plautus Scudder & Burgess, 1870; Proc. Bost. Soc. Nat. Hist. 13: 304; TL: Florida
    
    = ; [NL4A], #757a
  - Erynnis (Erynnides) juvenalis juvenalis; [NL4A], #757a
  - Erynnis juvenalis juvenalis; [iNaturalist]
  - Erynnis juvenalis plautus; ; [LepIndex]; [CoL]
  - Thanaos plautus; Dyar, 1903, Bull. U.S. nat. Mus. 52: 61
- E. j. clitus (Edwards, 1883) Arizona, New Mexico, Mexico
  - Thanaos clitus Edwards, 1882; Papilio 2 (9-10): 180; TL: Arizona
  - Thanaos maestus Godman, [1899]; Biol. centr.-amer., Lep. Rhop. 2: 457, 3: pl. 91, f. 18; TL: Arizona; Mexico, Pinal, Puebla
    
    = ; [NL4A], #757b
  - Erynnis (Erynnides) juvenalis clitus; [NL4A], #757b; [P&N23]
  - Erynnis clitus; [BOW]: pl. 21, f. 34 (text)
  - Erynnis juvenalis clitus; [NACL], #3947a; [iNaturalist]
  - Nisoniades moestus; Skinner, 1904, (missp.)
  - Thanaos clitus; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
Erynnis martialis (Scudder, 1869) NE.USA - Alabama. See [maps] Mottled Duskywing
- Nisoniades martialis Scudder, 1869; Trans. Chicago Acad. Sci. 1 (2): 355; TL: Iowa
- Thanaos quercus Butler, 1870; (nom. nud.)
  - = ; [NL4A], #758
- Nisoniades ausonius Lintner, 1872; 23 Rept. N.Y. Sta. Cab. Nat. Hist.: 166; TL: New York
  - = ; [NL4A], #758
- Erynnis (Erynnides) martialis; [NL4A], #758; [P&N23]
- Erynnis martialis; [NACL], #3954; [NAB], 753; [BOW]: pl. 21, f. 37; [Opler]; [MPG] ; [LepIndex] ; [CoL] ; [iNaturalist] ; [BAMONA]
- Thanaos ausonius; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
- Thanaos martialis; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
Larva on Ceanothus americanus, C. fendleri [NAB]
Erynnis meridianus Bell, 1927 California, Nevada, New Mexico, .... See [maps] Meridian Duskywing Southern Duskywing
- Erynnis meridianus Bell, 1927; J. N.Y. Ent. Soc. 35 (3): 261; TL: Arizona
- Erynnis (Erynnides) meridianus meridianus; [P&N23]
- Erynnis (Erynnides) meridianus; [NL4A], #759; [P&N23]
- Erynnis meridianus; [NACL], #3950; [NAB], 751 (note); [BOW]: pl. 21, f. 35 (text); [Opler]; [MPG] ; [LepIndex] ; [CoL] ; [iNaturalist] ; [BAMONA]
- E. m. meridianus Arizona, Mexico, ...
  - Erynnis (Erynnides) meridianus meridianus; [NL4A], #759a
  - Erynnis meridanus; Ferris, 1976, (missp.)
  - Erynnis meridianus meridianus; [iNaturalist]
- E. m. fieldi Burns, 1964 Mexico
  - Erynnis meridianus fieldi Burns, 1964; ; TL: Mexico, Michoacán
  - Erynnis (Erynnides) meridianus fieldi; [NL4A], #759b
  - Erynnis meridianus fieldi; [LepIndex]; [CoL] ; [iNaturalist]
Erynnis pacuvius (Lintner, 1878) W.USA, S.USA. See [maps] Pacuvius Duskywing Buckthorn Dusky Wing Dyar's Duskywing
- Nisoniades pacuvius Lintner, 1878; 30 Rept. N.Y. Mus. Nat. Hist.: 172; TL: New Mexico
- Erynnis (Erynnides) pacuvius pacuvius; [P&N23]
- Erynnis (Erynnides) pacuvius; [NL4A], #760; [P&N23]
- Erynnis pacuvius; [NACL], #3955; [NAB], 754; [BOW]: pl. 21, f. 33 (text); [BOC], 53; [Opler]; [MPG] ; [LepIndex] ; [CoL] ; [iNaturalist] ; [BAMONA]
- Thanaos pacuvius; Godman & Salvin, [1899], Biol. centr.-amer., Lep. Rhop. 2: 458, 3: pl. 91, f. 16-17; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
Larva on Ceanothus , C. fendleri [NAB]
- E. p. pacuvius Mexico, New Mexico
  - Erynnis (Erynnides) pacuvius pacuvius; [NL4A], #760a
  - Erynnis pacuvius pacuvius; [iNaturalist]
- E. p. lilius (Dyar, 1904) British Columbia, Washington
  - pacuvius lilius Dyar, 1904; ; TL: Kaslo, British Columbia
  - Erynnis (Erynnides) pacuvius lilius; [P&N23]
  - Erynnis pacuvius lilius; [NACL], #3955a; [BOC], 53; [LepIndex]; [CoL] ; [iNaturalist]
  Larva on Ceanothus velutinus [BOC]
- E. p. pernigra (Grinnell, 1905)
  - Erynnis (Erynnides) pacuvius pernigra; [P&N23]
  - Erynnis pacuvius perniger[sic]; [NACL], #3955b
  - Erynnis pacuvius pernigra; [iNaturalist]
  - ?Erynnis persius pernigra;
- E. p. callidus (Grinnell, 1904) California, Mexico
  - Thanaus callidus Grinnell, 1905; ; TL: California
  - Erynnis (Erynnides) pacuvius callidus; [NL4A], #760b; [P&N23]
  - Erynnis brizo callidus; [LepIndex]; [CoL]
  - Erynnis pacuvius callidus; [NACL], #3955c; [iNaturalist]
Erynnis propertius (Scudder & Burgess, 1870) from (British Columbia - Baja California) - Nevada. See [maps] Propertius Duskywing Western Oak Dusky Wing
- Nisoniades propertius Scudder & Burgess, 1870; Proc. Bost. Soc. Nat. Hist. 13: 298; TL: California
- Nisoniades tibullus Scudder & Burgess, 1870; Proc. Bost. Soc. Nat. Hist. 13: 298; TL: California
  - = ; [NL4A], #761
- ?Nisoniades[?] propertius Lintner, 1881; Papilio 1 (5): 71
- Erynnis (Erynnides) propertius; [NL4A], #761; [P&N23]
- Erynnis propertius; [NACL], #3949; [NAB], 751; [BOW]: pl. 21, f. 35 (text); [Opler]; [MPG] ; [LepIndex] ; [CoL] ; [iNaturalist] ; [BAMONA]
- Thanaos propertius Lintner, 1881; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
- Thanaos tibullus; Dyar, 1903, Bull. U.S. nat. Mus. 52: 61
Larva on Quercus agrifolia, Q. garryana [NAB]
Erynnis scudderi (Skinner, 1914) Mexico, Arizona. See [maps] Scudder's Duskywing
- Thanaos scudderi Skinner, 1914; ; TL: Arizona
- Erynnis (Erynnides) scudderi; [NL4A], #762; [P&N23]
- Erynnis scudderi; [NACL], #3951; [NAB], 753 (note); [Opler]; [MPG] ; [LepIndex] ; [CoL] ; [iNaturalist] ; [BAMONA]
Erynnis telemachus Burns, 1960 Arizona. See [maps] Rocky Mountain Duskywing
- Erynnis telemachus Burns, 1960; Wasmann J. Biol. 18 (1): 148; TL: Arizona
- Erynnis (Erynnides) telemachus; [NL4A], #763; [P&N23]
- Erynnis telemachus; [NACL], #3948; [NAB], 750; [Opler]; [MPG] ; [LepIndex] ; [CoL] ; [iNaturalist] ; [BAMONA]
Larva on Quercus gambelii [NAB]
Erynnis tristis (Boisduval, 1852) Arizona, New Mexico, Texas, Mexico - Colombia, CA. See [maps] Mournfull Duskywing
- Thanaos tristis Boisduval, 1852; Ann. Soc. ent. Fr. (2) 10 (2): 311; TL: California
- Erynnis (Erynnides) tristis tristis; [P&N23]
- Erynnis (Erynnides) tristis; [NL4A], #764; [P&N23]
- Erynnis tristis; [NACL], #3953; [NAB], 753; [Opler]; [MPG] ; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [BAMONA]
- Nisoniades tristis; Scudder & Burgess, 1870, Proc. Bost. Soc. Nat. Hist. 13: 303
- Thanaos tristis; Godman & Salvin, [1899], Biol. centr.-amer., Lep. Rhop. 2: 457, 3: pl. 91, f. 15; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
Larva on Quercus [NAB]
- E. t. tristis Mexico, California, ...
  - Erynnis (Erynnides) tristis tristis; [NL4A], #764a
  - Erynnis ristis; Warren, 2000, (missp.)
  - Erynnis tristis tristis; [iNaturalist]
  - Erynnis tristus; Ferris, 1977, (missp.)
  - Erynnis tritus; Ferris, 1977, (missp.)
- E. t. tatius (Edwards, 1882) Arizona, Mexico, Guatemala, Costa Rica, Panama - Colombia
  - Thanaos tatius Edwards, 1882; Papilio 2 (9-10): 179; TL: [Arizona]
  - Thanaos albomarginatus Godman, [1899]; Biol. centr.-amer., Lep. Rhop. 2: 458, 3: pl. 91, f. 19-23; TL: Mexico, Cordova; Guatemala, Chisoy Valley; Costa Rica, Irazu; Panama, Chiriqui; Colombia
    
    = ; [NL4A], #764c
  - Erynnis (Erynnides) tristis tatius; [NL4A], #764c; [P&N23]
  - Erynnis albomarginatus; [BOW]: pl. 21, f. 33
  - Erynnis tatitus; Ferris, 1977, (missp.)
  - Erynnis tristis tatius; [NACL], #3953a; [iNaturalist]
  - Nisoniades tacitus; Maynard, 1901, (missp.)
  - Thanaos tatius; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
- E. t. pattersoni Burns, 1964 Mexico(Baja California Sur)
  - Erynnis tristis pattersoni Burns, 1964; ; TL: Mexico(Baja California Sur)
  - Erynnis (Erynnides) tristis pattersoni; [NL4A], #764b
  - Erynnis tristis pattersoni; [LepIndex]; [CoL] ; [iNaturalist]
Erynnis zarucco (Lucas, 1857) Cuba, Texas, Florida. See [maps] Zarucco Duskywing
- Thanaos zarucco Lucas, 1857; ; TL: Cuba
- Nisoniades terentius Scudder & Burgess, 1870; Proc. Bost. Soc. Nat. Hist. 13: 292; TL: Florida
  - = ; [NL4A], #765
- Nisoniades ovidius Scudder & Burgess, 1870; ; TL: Florida
  - = ; [NL4A], #765
- Nisoniades naevius Lintner, 1881; Papilio 1 (5): 69; TL: Florida
  - = ; [NL4A], #765
- Antigonus diogenes Plötz, 1884; ; TL: Cuba
  - = ; [NL4A], #765
- Antigonus ferentius; Plötz, 1884, (missp.)
- Erynnis (Erynnides) zarucco; [NL4A], #765; [P&N23]
- Erynnis zarucco; [NACL], #3956; [NAB], 755; [BOW]: pl. 21, f. 35 (text); [Opler]; [MPG] ; [LepIndex] ; [CoL] ; [iNaturalist] ; [BAMONA]
- Erynnis zaruco; Silva et al., 1968, (missp.)
- Nisoniades jaracco; Wolcott, 1936, (missp.)
- Nisoniades jaruco; Gundlach, 1881, (missp.)
- Thanaos naevius; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
- Thanaos terentius; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
- jarucco; Beattie, 1976, (missp.)
Larva on Robinia [NAB]

Subgenus

Erynnis lucilius (Scudder & Burgess, 1870) S.USA - Mexico. See [maps] Columbine Duskywing
- lucilius (Scudder & Burgess, 1870) (Nisoniades); Proc. Bost. Soc. Nat. Hist. 13: 287
- ?lucilius Lintner, 1872; 23 Rept. N.Y. Sta. Cab. Nat. Hist.: 162
- Erynnis (Erynnides) lucilius; [P&N23]
- Erynnis lucilius; [NACL], #3958; [NAB], 756; [BOW]: pl. 21, f. 38 (text); [Opler]; [MPG] ; [LepIndex] ; [CoL] ; [iNaturalist] ; [BAMONA]
- ?Thanaos lucilius Lintner, 1872; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
Larva on Aquilegia canadensis [NAB]
Erynnis baptisiae (Forbes, 1936) E.USA. See [maps] Wild Indigo Duskywing
- Erynnis (Erynnides) baptisiae; [P&N23]
- Erynnis baptisiae; [NACL], #3959; [NAB], 757; [BOW]: pl. 21, f. 38 (text); [Opler]; [MPG] ; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [BAMONA]
Larva on Rubus chamaemorus [NAB]
Erynnis persius (Scudder, 1863) USA, Canada. See [maps] Hairy Dusky Wing Persius Duskywing
- persius Scudder, 1863; Proc. Essex Inst. 3: 170
- Erynnis (Erynnides) persius persius; [P&N23]
- Erynnis (Erynnides) persius; [P&N23]
- Erynnis persius; [NACL], #3961; [NAB], 758; [BOW]: pl. 21, f. 38; [BOC], 55; [Opler]; [MPG] ; [LepIndex] ; [CoL] ; [iNaturalist] ; [BAMONA]
- Thanaos persius; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
Larva on Lupinus , Thermopsis [NAB], Lupinus latifolius, L. sericeus, Thermopsis , Astragalus , Lotus [BOC]
- E. p. persius
  - Erynnis persius persius; [iNaturalist]
- E. p. borealis (Cary, 1906)
  - Thanaus propertius var. borealis Cary, 1906; Proc. U.S. Nat. Mus. 31 (1488): 455; TL: N.Nahanni R., Mckenzie Dist., NW Terr.
  - Erynnis (Erynnides) persius borealis; [P&N23]
  - Erynnis persius borealis; [NACL], #3961a; [BOC], 55; [LepIndex]; [CoL] ; [iNaturalist]
- E. p. avinoffi (Holland, 1930)
  - Erynnis (Erynnides) persius avinoffi; [P&N23]
  - Erynnis persius avinoffi; [NACL], #3961b; [iNaturalist]
- E. p. fredericki Freeman, 1943
  - Erynnis persius fredericki Freeman, 1943; Ent. News 54 (3): (72-77) ; TL: near Lead, Spearfish Canyon, [Lawrence Co.,] South Dakota
  - Erynnis (Erynnides) persius fredericki; [P&N23]
  - Erynnis persius frederecki[sic]; [BOC], 55
  - Erynnis persius fredericki; [NACL], #3961c; [iNaturalist]

Unmatched external taxa

Erynnis burgessi; [BOA]
Erynnis burgessi burgessi; [BOA]
Erynnis burgessi muelleri; [BOA]
Erynnis (Gesta) invisus; [P&N23]
Erynnis invisus; [MPG] ; [iNaturalist]
Erynnis lacustra; [BOA]
Erynnis lassauxi; [iNaturalist]
Erynnis somnus; [BOA]

Unknown or unplaced taxa

Thanaos tages f. posticeprivata Stauder, 1924; Mitt. Münch. Ent. Ges. 14 (1-5): 66

26.6.2023 (24)

References:

[BAMONA] Opler, Pavulaan, Stanford & Pogue
Butterflies and Moths of North America; Mountain Prairie Information Node
[BCA] electronic
BIOLOGIA CENTRALI-AMERICANA
[BOA] Interactive Listing of American Butterflies
Listado Interactivo de la Mariposas Americanas; Butterflies of America
[CiNii]
http://ci.nii.ac.jp/
[CoL] Catalogue of Life
Bánki, O., Roskov, Y., Döring, M., Ower, G., Hernández Robles, D. R., Plata Corredor, C. A., Stjernegaard Jeppesen, T., Örn, A., Vandepitte, L., Hobern, D., Schalk, P., DeWalt, R. E., Ma, K., Miller, J., Orrell, T., Aalbu, R., Abbott, J., Adlard, R., Aedo, C., et al. (2024). Catalogue of Life Checklist (Version 2024-03-26); https://doi.org/10.48580/dfz8d
[EVERSMANNIA]
Eversmannia contents
[HBN] Francis Hemming, 1937
HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[Heiner Ziegler]
Switzerland / Schweiz; www.pieris.ch
[LAMAS-NEO] Gerardo Lamas
Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)
[LEPIFORUM]
Lepiforum; Bestimmung von Schmetterlingen (Lepidoptera) und ihren Präimaginalstadien
[LepIndex] Global Lepidoptera Index
Beccaloni, G., Scoble, M., Kitching, I., Simonsen, T., Robinson, G., Pitkin, B., Hine, A., Lyal, C., Ollerenshaw, J., Wing, P., & Hobern, D. (2024). Global Lepidoptera Index. In O. Bánki, Y. Roskov, M. Döring, G. Ower, D. R. Hernández Robles, C. A. Plata Corredor, T. Stjernegaard Jeppesen, A. Örn, L. Vandepitte, D. Hobern, P. Schalk, R. E. DeWalt, K. Ma, J. Miller, T. Orrell, R. Aalbu, J. Abbott, R. Adlard, C. Aedo, et al., Catalogue of Life Checklist (1.1.24.106); https://doi.org/10.48580/dg4lg-49xk
[MBE] Moths and Butteflies of Europe
Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[MPG]
North American Moth Photographers Group
[NHM-BMW] Natural History Museum
Butterflies and Moths of the World; Generic Names and their Type-species
[Oleg Kosterin] Institue of Cytology & Genetics
Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[Opler] Opler & Warren, 2003
Butterflies of North America. 2. Scientific Names List for Butterfly Species of North America, north of Mexico.
[P&N23] Pohl & Nanz, 2023
Annotated Taxonomic Checklist of the Lepidoptera of North America, north of Mexico
[SVF]
Svenska fjärilar; Naturhistoriska riksmuseet; (Sweden)
[SZMN] Lepidoptera
Siberian Zoological Museum
[Simon Coombes]
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[USFCA]
[ZOBODAT] Oberösterreichische Landesmuseum
OÖ. Landesmuseum; ZOBODAT - Literatur
[iNaturalist]
iNaturalist
[maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.

Some related literature:

Bell, 1927
Description of a new species of Erynnis (Thanaos Auct.) (Lepidoptera-Rhopalocera-Hesperiidae) J. N.Y. Ent. Soc. 35 (3) : 261-263
Benyamini, Ugarte, Shapiro, Mielke, Pyrcz & Bálint, 2014
An updated list of the butterflies of Chile (Lepidoptera, Papilionoidea and Hesperioidea) including distribution, flight period and conservation status. Part I, Comprising the families: Papilionidae, Pieridae, Nympalidae (in part) and Hesperiidae. Describing a new species of Hypsochila (Pieridae) and a new subspecies of Yramea modesta (Nymphalidae) Bol. Mus. Nac. Hist. Nat. Chile 63 : 9-31
Boisduval, [1834]
Icones historique des Lépidoptères nouveaux ou peu connus. Collection, avec Figures coloritées, des Papillons d'Europe nouvellement découverts; ouvrage formant le complément de tous les auteurs iconographes Icon. hist. Lépid. Europe 1 (1-2): 1-24, pl. 1-4 (1832), (3-4): 25-40, pl. 5-8 (1832), (5-6): 41-56, pl. 9-12 (1832), (7-8): 57-72, pl. 13-16 (1832), (9-10): 73-104, pl. 17-20 (1833), (11-12): 105-120, pl. 21-24 (1833), (13-14): 121-144, pl. 25-28 (1833), (15-16): 145-160, pl. 29-32 (1833), (17-18): 161-184, pl. 33-36 (1833), (19-20): 185-208, pl. 37-40 (1834), (21-22): 209-224, pl. 41-44 (1834), (23-24): 225-248, pl. 45-47 (1834), (25-28): 249-252 (1834)
Boisduval, 1852
Lépidoptères de la Californie Ann. Soc. ent. Fr. (2) 10 (2) : 275-324
Boisduval & Le Conte, [1834]
Histoire générale et iconographie des lépidoptères et des chenilles de l'Amérique septentrionale Hist. Lép. Am. Sept. (1): 1-16. pl. 1-3 (1829), (2): 17-24, pl. 4-6 (1829), (3/4): 25-40, pl. 7-12 ([1830]), (5/6): 41-56, pl. 13-18 ([1830]), (7/8): 57-80, pl. 19-24 ([1830]), (9/10): 81-100, pl. 25-30 (1833), (11-22): 101-196, pl. 31-65 ([1835]), (23-26): 197-228, pl. 66-78 ([1837])
Bremer, 1861
Neue Lepidopteren aus Ost-Sibirien und dem Amur-Lande gesammelt von Radde und Maack, bescrieben von Otto Bremer Bull. Acad. Imp. Sci. St. Petersb. 3 : 461-496, Mélanges biol. St.-Pétersb. 3 (5-6): 538-589
Burns, 1960
A new species of oak-eating Erynnis (Lepidoptera: Hesperiidae) Wasmann J. Biol. 18 (1) : 147-160, f. 1-7
Butler, 1866
A list of the diurnal Lepidoptera recently collected by Mr. Whitely in Hakodadi (North Japan) J. Linn. Soc. Zool. Lond. 9 (34) : 50-58
Bytinski-Salz & Brandt, 1937
New Lepidoptera from Iran Ent. Rec. J. Var. (Suppl.) 49 : 1-15, : 9-15
Cary, 1906
On the Diurnal Lepidoptera of the Athabaska and Mackenzie Region, British America Proc. U.S. Nat. Mus. 31 (1488) : 425-457
[MRS]; Chou, 1994
Monografia Rhopalocerorum Sinensium 1-2 Mon. Rhop. Sinesium : 1-854
Dunning & Pickard, 1858
An Accentuated List of the British Lepidoptera Accent. List Brit. Lep.: : 1-118
Dyar, 1903
A List of North American Lepidoptera and Key to the Literature of this Order of Insects Bull. U.S. nat. Mus. 52 : 1-723
Edwards, 1882
Two new species of Thanaos Papilio 2 (9-10) : 179-180
Evans, 1949
A catalogue of the Hesperiidae from Europe, Asia and Australia in the British Museum Cat. Hesp. Europe Asia Australia Brit. Mus. : 1-502, pl. 1-53
Fabricius, 1793
Entomologia Systematica emendata et aucta. Ent. Syst. 3 (1) : 1-487
Fabricius, 1807
Die neueste Gattungs-Eintheilung der Schmetterlinge aus den Linnéischen Gattungen Papilio und Sphinx Mag. f. Insektenk. 6 : 277-295
Freeman, 1943
New Hesperioidea, with Notes on Some Others from the United States (Lepidotera, Rhopalocera) Ent. News 54 (3) : 72-77
Godman & Salvin, [1899]; Godman in Godman & Salvin, [1899]
Biologia Centrali-Americana. Rhopalocera. (1887-1901) Biol. centr.-amer., Lep. Rhop. 2: 1-741 (?782), 3: pl. 1-112
Grum-Grshimailo, 1891
Lepidoptera nova in Asia centrali novissime lecta et descripta Horae Soc. ent. Ross. 25 (3-4) : 445-465
[H&R]; Higgins & Riley, 1970
A Field Guide to the Butterflies of Britain and Europe
[NACL]; Hodges, 1983
Check List of the Lepidoptera of America North of Mexico Check List Lep. Am. N of Mexico
Hübner, 1822
Systematisch-alphabetisches Verzeichniss aller bisher bey den Fürbildungen zur Sammlung europäischer Schmetterlinge angegebenen Gattungsbenennungen; mit Vormerkung auch augsburgischer Gattungen, von Jacob Hübner Syst.-alph. Verz. : I-III, IV-VI, 1-81
Korb & Bolshakov, 2011
[A catalogue of butterflies (Lepidoptera: Papilioformes) of the former USSR. Second edition, reformatted and updated] (in Russian) Eversmannia Suppl. 2 : 1-123
Kudrna, 1974
An annotated List of Japanese Butterflies Atalanta 5 : 92-120
[NL4A]; Lamas, 2004
Checklist: Part 4A, Hesperioidea - Papilionoidea Atlas of Neotropical Lepidoptera
Leech, 1891
New Species of Rhopalocera from Western China Entomologist 24 (Suppl.) : 57-61, 66-68
[RFEL]; Leley, 2016
Annotated catalogue of the insects of Russian Far East. Volume II. Lepidoptera Cat. ins. Russian Far East. 2 : 1-812
[BOW]; Lewis, 1974
Butterflies of the World
Linnaeus, 1758
Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338, : 339-824
Lintner, 1881
On some species of Nisoniades Papilio 1 (5) : 69-74
Mabille, 1883
Description d'hespériens Bull. Soc. Ent. Belg. 1883 : li-lxxviii
Murayama & Yoshisaka, 1959
Die neu aus Formosa entdeckte Art Erynnis montanus BREMER [in Japanese] Tyô to Ga 10 (1): 14
Nordmann, 1851
Neue Schmetterlinge Russlands Bull. Soc. imp. Nat. Moscou 24 : 439-446, pl. 11-12
Oken, 1815
Okens Lehrbuch der Zoologie Lehrbuch Naturgesch. 3 (Zool.) (1) : 1-850
[NAB]; Pyle, 1981
Field Guide to North American Butterflies, National Audubon Society
[BOC]; Pyle, 2002
The Butterflies of Cascadia
Schrank, 1801
Fauna Boica. Durchgedachte Geschichte der in Baiern einheimischen und zahmen Thiere Fauna Boica 2 (1) : 1-374
Scudder, 1863
A List of the Butterflies of New England Proc. Essex Inst. 3 : 161-179
Stauder, 1924
Neue Palaearktenformen. II Mitt. Münch. Ent. Ges. 14 (1-5) : 59-66
Tutt, 1906
A natural history of the British Lepidoptera. A text-book for students and collectors Nat. Hist. Br. Lepid. 8 : 1-479, pl. 1-20
[BRU]; Tuzov, Bogdanov, Devyatkin, Kaabak, Korolev, Murzin, Samodurov, Tarasov, 1997
Guide to the Butterflies of Russia and adjacent territories: Hesperiidae, Papilionidae, Pieridae, Satyridae Butts. Russia adj. terr. 1
Yakovlev, 2012
Checklist of Butterflies (Papilionoidea) of the Mongolian Altai Mountains, including descriptions of new taxa Nota lepid. 35 (1) : 51-96

If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.

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