Euphilotes

Euphilotes Mattoni, [1978]

Euphilotes Mattoni, [1978]; J. Res. Lepid. 16 (4): 224; TS: Lycaena enoptes Boisduval

Euphilotes; [P&N23]; [CoL] ; [iNaturalist] ; [NHM-BMW]
Euphilotes (Section Glaucopsyche); Ugelvig, Vila, Pierce & Nash, 2011, Molec. Phyl. Evol. 61: 238
Euphilotes (Polyommatinae); [NL4A], 139
Euphilotes (Polyommatini); [Opler]; Pelham, 2008, J. Res. Lepid. 40: 246
Euphilotes (Scolitantidina); Stradomsky, 2016, Caucasian Ent. Bull. 12 (1): 151

BATTOIDES-complex

Battoides; Pratt & Emmel, 1998, Syst. W. N. Am. Butts. (16): 216; Pelham, 2008, J. Res. Lepid. 40: 248

Euphilotes battoides (Behr, 1867) -. See [maps] Square-spotted Blue Buckwheat Blue
- Lycaena battoides Behr, 1867; Proc. Calif. Acad. Sci. 3 (4): 282; TL: California
- Euphilotes battoides; [NACL], #4366; [NAB]; [BNA]: 401, pl. 37, f. 399; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 246; [MPG] ; [P&N23]; [CoL] ; [iNaturalist] ; [iNaturalist] ; [BOA] ; [BOA] ; [BAMONA]
- Philotes battoides; [BOW]: pl. 20, f. 13; [iNaturalist]
- Rusticus battoides; Dyar, 1903, Bull. U.S. nat. Mus. 52: 44
identification uncertain 1200x1261 (~192Kb) underside USA: Washington, Chelan Co., LepSoc 2010 Field Trip (Eagle Creek ?), 11.7.2010, Photo © Markku Savela

Larva on Eriogonum [NAB]
- E. b. battoides California
  - Euphilotes battoides battoides; [BNA]; Pelham, 2008, J. Res. Lepid. 40: 247; [P&N23]; [iNaturalist] ; [BOA]
  Larva on Eriogonum lobbii var. lobbii, E. incanum, E. polypodum [BNA]
- E. b. allyni (Shields, 1975) California
  - Shijiamiaeoides battoides allyni Shields, 1975; Bull. Allyn Mus. 28: 10; TL: California, Los Angeles Co., El Segundo
  - Euphilotes allyni; [MPG] ; [P&N23]
  - Euphilotes battoides allyni; [NACL], #4366i; [BNA]; Pelham, 2008, J. Res. Lepid. 40: 247; [iNaturalist]
  Larva on Eriogonum parvifolium var. parvifolium, E. cinereum, E. fasciculatum var. fasciculatum [BNA]
- E. b. argocyanea Pratt & Emmel, 1998 California
  - Euphilotes battoides argocyanea Pratt & Emmel, 1998; Syst. W. N. Am. Butts. (16): 217; TL: California, Inyo Co., E end of Nelson Range: slope 0.3km W of Junction of Saline Valley Road and Hunter Mt Road, 1800-1900m
  - Euphilotes battoides argocyanea; Pelham, 2008, J. Res. Lepid. 40: 247; [P&N23]; [iNaturalist] ; [BOA]
- E. b. mazourka Pratt & Emmel, 1998 California
  - Euphilotes battoides mazourka Pratt & Emmel, 1998; Syst. W. N. Am. Butts. (16): 218; TL: California, Inyo Co., Inyo Mtns, NW edge of Badger Flat
  - Euphilotes battoides mazourka; Pelham, 2008, J. Res. Lepid. 40: 247; [P&N23]; [iNaturalist] ; [BOA]
- E. b. panamintensis Pratt & Emmel, 1998 California
  - Euphilotes battoides panamintensis Pratt & Emmel, 1998; Syst. W. N. Am. Butts. (16): 219; TL: California, Inyo Co., Panamint Range, Wildrose Cyn., 2100m
  - Euphilotes battoides panamintensis; Pelham, 2008, J. Res. Lepid. 40: 247; [P&N23]; [iNaturalist] ; [BOA]
- E. b. vernalis Pratt & Emmel, 1998 California
  - Euphilotes battoides vernalis Pratt & Emmel, 1998; Syst. W. N. Am. Butts. (16): 221; TL: California, San Bernardino Co., Coxey Meadow, 1600m
  - Euphilotes battoides vernalis; Pelham, 2008, J. Res. Lepid. 40: 247; [iNaturalist]
  - Euphilotes baueri vernalis; [P&N23]
- E. b. fusimaculata Austin, 1998 Nevada
  - Euphilotes battoides fusimaculata Austin, 1998; Syst. W. N. Am. Butts. (45): 551; TL: Nevada, Nye Co., Toiyable Mtns, Peavine Creek Campground, 2012m
  - Euphilotes battoides fusimaculata; Pelham, 2008, J. Res. Lepid. 40: 247; [P&N23]; [iNaturalist] ; [BOA]
- E. b. anasazi Scott, 1998 New Mexico
  - Euphilotes battoides anasazi Scott, 1998; Papilio (n.s.) 11: 5; TL: Aztec, San Juan Co., New Mexico
  - Euphilotes battoides anasazi; Pelham, 2008, J. Res. Lepid. 40: 247; [iNaturalist] ; [BOA]
  - Euphilotes ellisii anasazi; [P&N23]
Euphilotes glaucon (Edwards, 1871) British Columbia - NE.California, S.Idaho. See [maps]
- Lycaena glaucon Edwards, 1871; Trans. amer. ent. Soc. 3 (3/4): 210; TL: Nevada [Storey Co.]
- Euphilotes glaucon; [BNA]; Pelham, 2008, J. Res. Lepid. 40: 247; [MPG] ; [P&N23]; [iNaturalist] ; [BOA] ; [BOA] ; [BOA] ; [BOA] ; [BAMONA]
- Philotes glaucon; [BOW]: pl. 20, f. 14 (text only)
- Rusticus glaucon; Dyar, 1903, Bull. U.S. nat. Mus. 52: 44
Larva on Eriogonum umbellatum, Eriogonum ovalifolium var. nivale, E. hreacleiodes, E. sphaerocephalum var. halimioides, E. flavum var. piperi [BNA]
- E. g. glaucon Nevada
  - Euphilotes battoides glaucon; [NACL], #4366b
  - Euphilotes glaucon glaucon; Pelham, 2008, J. Res. Lepid. 40: 247; [P&N23]; [BOA]
- E. g. oregonensis (Barnes & McDunnough, 1917) S.Oregon
  - Philotes glaucon oregonensis Barnes & McDunnough, 1917; Contr. nat. Hist. Lep. N. Am. 3 (4): 214; TL: Crater Lake, Oregon
    
    = Euphilotes intermedia; [Opler]
  - Euphilotes battoides oregonensis; [NACL], #4366a; [CoL]
  - Euphilotes glaucon oregonensis; [BNA]; Pelham, 2008, J. Res. Lepid. 40: 247; [P&N23]; [iNaturalist] ; [BOA]
  Larva on Eriogonum marifolium [BNA]
- E. g. intermedia (Barnes & McDunnough, 1917) California
  - Philotes glaucon intermedia Barnes & McDunnough, 1917; Contr. nat. Hist. Lep. N. Am. 3 (4): 214, pl. 16, f. 4-6; TL: Shasta Co., California
  - Philotes glaucon intermedia tr. f. malcolmi Gunder, 1927; Can. Ent. 59 (12): 282; TL: Amer. River, Placer Co., California
    
    = ; Pelham, 2008, J. Res. Lepid. 40: 248
  - Euphilotes battoides intermedia; [NACL], #4366d
  - Euphilotes battoides malcolmi; [CoL]
  - Euphilotes glaucon intermedia; [BNA]; Pelham, 2008, J. Res. Lepid. 40: 248; [P&N23]; [iNaturalist] ; [BOA]
  - Euphilotes intermedia; [Opler]; [BAMONA]
  - Philotes intermedia; [CoL]
  Larva on Eriogonum incanum, E. marifolium [BNA]
- E. g. comstocki (Shields, 1975) California
  - Shijimiaeoides battoides comstocki Shields, 1975; Bull. Allyn Mus. 28: 12; TL: California, Kern Co., Tehachapi
  - Euphilotes battoides comstocki; [NACL], #4366e
  - Euphilotes glaucon comstocki; [BNA]; Pelham, 2008, J. Res. Lepid. 40: 248; [P&N23]; [iNaturalist] ; [BOA]
  Larva on Eriogonum umbellatum [BNA]
- E. g. australoglaucon Pratt & Emmel, 1998 California
  - Euphilotes battoides australoglaucon Pratt & Emmel, 1998; Syst. W. N. Am. Butts. (16): 216; TL: California, Inyo Co., E slope Sierra Nevada, Independence Creek, 6300'
  - Euphilotes glaucon australoglaucon; Pelham, 2008, J. Res. Lepid. 40: 247; [P&N23]; [BOA]
Euphilotes centralis (Barnes & McDunnough, 1917) S.Colorado - N.New Mexico. See [maps]
- Philotes glaucon centralis Barnes & McDunnough, 1917; Contr. nat. Hist. Lep. N. Am. 3 (4): 215, pl. 16, f. 7-8; TL: Salida, Colorado
- Euphilotes battoides centralis; [NACL], #4366f; [CoL] ; [iNaturalist]
- Euphilotes centralis; Pelham, 2008, J. Res. Lepid. 40: 248; [MPG] ; [P&N23]; [iNaturalist] ; [BOA] ; [BAMONA]
- E. c. centralis Colorado
  - Euphilotes centralis centralis; Pelham, 2008, J. Res. Lepid. 40: 248; [P&N23]; [iNaturalist] ; [BOA]
- E. c. hadrochilus Pratt & Emmel, 1998 California
  - Euphilotes centralis hadrochilus Pratt & Emmel, 1998; Syst. W. N. Am. Butts. (16): 222; TL: California, Inyo Co., 6km S of Westgard Pass, 2000m
  - Euphilotes centralis hadrochilus; Pelham, 2008, J. Res. Lepid. 40: 248; [P&N23]; [iNaturalist] ; [BOA]
Euphilotes bernardino (Barnes & McDunnough, 1916) -. See [maps]
- Philotes battoides bernardino Barnes & McDunnough, 1916; Contr. nat. Hist. Lep. N. Am. 3 (2): 116, pl. 11, f. 9, 11, 13; TL: California
- Euphilotes battoides bernardino; [NACL], #4366h
- Euphilotes bernardino; [Opler]; [NL4A], #1091; [BNA]; Pelham, 2008, J. Res. Lepid. 40: 248; [MPG] ; [P&N23]; [iNaturalist] ; [BOA] ; [BAMONA]
- Philotes bernardino; [CoL]
Larva on Eriogonum fasciculatum, E. heermannii [BNA]
- E. b. bernardino California
  - Philotes battoides var. bernardino ab. ♀ baldyensis Gunder, 1925; Ent. News 36: 3, pl. 1, f. M; TL: California
    
    = ; [NL4A], #1091a; Pelham, 2008, J. Res. Lepid. 40: 248
  - Euphilotes bernardino bernardino; [NL4A], #1091a; Pelham, 2008, J. Res. Lepid. 40: 248; [P&N23]; [iNaturalist] ; [BOA]
- E. b. martini (Mattoni, 1954) California - S.Arizona
  - Philotes battoides martini Mattoni, 1954; Bull. Sth. Calif. Acad. Sci. 53 (3): 157, 159, pl. 43, f. 3-4; TL: Oatman, Mohave Co., Arizona
  - Euphilotes battoides martini; [NACL], #4366j; [CoL]
  - Euphilotes bernardino martini; [BNA]; [NL4A], #1091c; Pelham, 2008, J. Res. Lepid. 40: 248; [P&N23]; [iNaturalist] ; [BOA]
  - Philotes battoides martini; [iNaturalist]
  Larva on Eriogonum fasciculatum ssp. polifolium [BNA]
- E. b. garthi Mattoni, 1989 Baja California
  - Euphilotes bernardino garthi Mattoni, 1989; J. Res. Lepid. 27 (3-4): 180; TL: Mexico, Baja California
  - Euphilotes bernardino garthi; [NL4A], #1091b; [iNaturalist] ; [BOA]
- E. b. inyomontana Pratt & Emmel, 1998 California
  - Euphilotes bernardino inyomontana Pratt & Emmel, 1998; Syst. W. N. Am. Butts. (16): 223; TL: California, Inyo Co., Panamint Range, S end, ridge just above Wood Canyon, 1500-1700m, 1.3km NNW of Manly Peak
  - Euphilotes bernardino inyomontana; Pelham, 2008, J. Res. Lepid. 40: 248; [P&N23]; [iNaturalist] ; [BOA]
- E. b. minuta Austin, 1998 Nevada
  - Euphilotes bernardino minuta Austin, 1998; Syst. W. N. Am. Butts. (45): 549; TL: Nevada, White Pine Co., Spring Valley, Baking Powder Flat, 1 mile S of Blind Spring, 1762m
  - Euphilotes bernardino minuta; Pelham, 2008, J. Res. Lepid. 40: 248; [P&N23]; [iNaturalist] ; [BOA]
Euphilotes ellisii (Shields, 1975) -. See [maps]
- Shijimjiaeoides battoides ellisii Shields, 1975; Bull. Allyn Mus. 28: 12; TL: Colorado, Mesa Co., W. Creek, Unaweep Canyon, 5400', 6mi E of Gateway
- Euphilotes battoides ellisii; [NACL], #4366g
- Euphilotes ellisi; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 248; [BAMONA]
- Euphilotes ellisii ellisi; [iNaturalist]
- Euphilotes ellisii euromajavensis; [iNaturalist]
- Euphilotes ellisii; [MPG] ; [P&N23]; [iNaturalist] ; [BOA]
- E. e. ellisii Colorado
  - Euphilotes ellisii ellisii; Pelham, 2008, J. Res. Lepid. 40: 248; [P&N23]; [iNaturalist] ; [BOA]
- E. e. euromojavensis Pratt & Emmel, 1998 California
  - Euphilotes ellisii euromojavensis Pratt & Emmel, 1998; Syst. W. N. Am. Butts. (16): 225; TL: California, San Bernardino Co., Mid Hills, N facing slope of Macedonia Canyon, 1400-1500m
  - Euphilotes ellisii euromojavensis; Pelham, 2008, J. Res. Lepid. 40: 249; [P&N23]; [BOA]
- E. e. avawatz (Pratt & Emmel, 1998) California
  - Eupholites ellisii avawatz Pratt & Emmel, 1998; Syst. W. N. Am. Butts. (16): 226; TL: California, San Bernardino Co., NE Mojave Desert, Avawatz Mtns, 5km W of Old Mormon Spring, 1100m
  - Euphilotes ellisii avawatz; [P&N23]; [iNaturalist] ; [BOA]
  - Eupholites ellisii avawatz; Pelham, 2008, J. Res. Lepid. 40: 249
- E. e. basinensis Austin, 1998 Nevada
  - Euphilotes ellisii basinensis Austin, 1998; Syst. W. N. Am. Butts. (45): 551; TL: Nevada, Lyon Co., Pine Nut Mtns, Eldorado Canyon, 3-4mil SE of Dayton, 1524m
  - Euphilotes ellisii basinensis; Pelham, 2008, J. Res. Lepid. 40: 249; [P&N23]; [iNaturalist] ; [BOA]
Euphilotes baueri (Shields, 1975) California. See [maps]
- Shijimiaeoides battoides baueri Shields, 1975; Bull. Allyn Mus. 28: 15; TL: California, Inyo Co., W side Gilbert Pass, 6200'
- Euphilotes battoides baueri; [NACL], #4366c
- Euphilotes baueri borealis; [P&N23]; [iNaturalist]
- Euphilotes baueri shoshone; [P&N23]; [iNaturalist]
- Euphilotes baueri; [Opler]; [BNA]; Pelham, 2008, J. Res. Lepid. 40: 249; [MPG] ; [P&N23]; [iNaturalist] ; [BOA] ; [BOA] ; [BAMONA]
Larva on Eriogonum ovalifolium, Eriogonum shockleyi ssp. longilobum [BNA]
- E. b. baueri California
  - Euphilotes baueri baueri; Pelham, 2008, J. Res. Lepid. 40: 249; [P&N23]; [iNaturalist] ; [BOA]
- E. b. orientis Austin, 1998 Nevada
  - Euphilotes baueri orientis Austin, 1998; Syst. W. N. Am. Butts. (45): 550; TL: Nevada, Lincoln Co., Bristol Mtns, Hwy 92, 12.5km road miles N of Pioche, 1826m
  - Euphilotes baueri orientis; Pelham, 2008, J. Res. Lepid. 40: 249; [P&N23]; [iNaturalist] ; [BOA]

ENOPTES-complex

Enoptes; Pratt & Emmel, 1998, Syst. W. N. Am. Butts. (16): 209; Pelham, 2008, J. Res. Lepid. 40: 249

Euphilotes mojave (Watson & Comstock, 1920) -. See [maps]
- Philotes eunoptes mojave Watson & Comstock, 1920; Bull. Amer. Mus. 42 (10): 455; TL: Mojave Desert, California
- Euphilotes mojave virginensis; [P&N23]; [iNaturalist]
- Euphilotes mojave; [Opler]; [BNA]; Pelham, 2008, J. Res. Lepid. 40: 249; [MPG] ; [P&N23]; [CoL] ; [iNaturalist] ; [BOA] ; [BAMONA]
- mohave[sic]; auth
Larva on Eriogonum nudum, Eriogonum pusillum, E. kennedyi [BNA]
- E. m. mojave California
  - Euphilotes enoptes mojave; [NACL], #4367e
  - Euphilotes mojave mojave; Pelham, 2008, J. Res. Lepid. 40: 249; [P&N23]; [iNaturalist] ; [BOA]
- E. m. virginiensis Austin, 1998 Nevada
  - Euphilotes mojave virginiensis Austin, 1998; Syst. W. N. Am. Butts. (45): 556; TL: Nevada, Clark Co., Virgin Mtns, Cabin Canyon, 1341m
  - Euphilotes mojave virginiensis; Pelham, 2008, J. Res. Lepid. 40: 249
Euphilotes enoptes (Boisduval, 1852) W.USA (Pacific states). See [maps] Dotted Blue
- Lycaena enoptes Boisduval, 1852; Ann. Soc. ent. Fr. (2) 10 (2): 298
- Euphilotes enoptes; [NACL], #4367; [BNA]: 403, pl. 37, f. 400; [NAB], 498; [Opler]; [NL4A], #1092; Pelham, 2008, J. Res. Lepid. 40: 249; [MPG] ; [P&N23]; [CoL] ; [iNaturalist] ; [BOA] ; [BOA] ; [BOA] ; [BOA] ; [BOA] ; [BAMONA]
- Philotes enoptes; [BOW]: pl. 20, f. 14
- Rusticus enoptes; Dyar, 1903, Bull. U.S. nat. Mus. 52: 44
identification uncertain 600x457 (~46Kb) ♂ underside USA: Robinson Canyon Rd, (47°01'00N 120°43'40W), Kittitas Co., WA, 26.7.1999, Photo © Markku Savela

Larva on Eriogonum [NAB]
- E. e. enoptes SW.Oregon - California
  - Euphilotes enoptes enoptes; [BNA]; Pelham, 2008, J. Res. Lepid. 40: 249; [P&N23]; [iNaturalist] ; [BOA]
  Larva on Eriogonum nudum, Eriogonum wrightii var. subcaposum, E. roseum, E. saxatile, E. davidsoni [BNA]
- E. e. dammersi (Comstock & Henne, 1933) S.Nevada, California, Arizona
  - Philotes enoptes dammersi Comstock & Henne, 1933; Bull. Sth. Calif. Acad. Sci. 32 (1): 24; TL: Snow Creek, Riverside Co., California
  - Euphilotes dammersi dammersi; [P&N23]
  - Euphilotes dammersi; [MPG] ; [P&N23]
  - Euphilotes enoptes dammersi; [NACL], #4367a; [NL4A], #1092b; [BNA]; Pelham, 2008, J. Res. Lepid. 40: 250; [CoL] ; [iNaturalist] ; [BOA]
  Larva on Eriogonum nudum var. pauciflorum, E. elongatum var. elongatum, E. wrightii [BNA]
- E. e. smithi (Mattoni, 1954) California
  - Philotes enoptes smithi Mattoni, 1954; Bull. Sth. Calif. Acad. Sci. 53 (3): 160, pl. 43, f. 8-9; TL: Burns Creek, State Hwy 1, Monterey Co., California
  - Euphilotes enoptes smithi; [NACL], #4367d; [BNA]; Pelham, 2008, J. Res. Lepid. 40: 250; [P&N23]; [CoL] ; [iNaturalist] ; [BOA]
  Larva on Eriogonum latifolium, E. parvifolium [BNA]
- E. e. bayensis (Langston, 1964) California
  - Philotes enoptes bayensis Langston, 1964; J. Lep. Soc. 17 (4): 208; TL: China Camp, nr Point San Pedro, Marin Co., California
  - Euphilotes enoptes bayensis; [NACL], #4367h; [BNA]; Pelham, 2008, J. Res. Lepid. 40: 250; [P&N23]; [CoL] ; [iNaturalist] ; [BOA]
  Larva on Eriogonum nudum [BNA]
- E. e. tildeni (Langston, 1964) California
  - Philotes enoptes tildeni Langston, 1964; J. Lep. Soc. 17 (4): 212; TL: Del Puerto Canyon, 22 mi. W Patterson, Stanislaus Co., California
  - Euphilotes enoptes tildeni; [NACL], #4367b; [BNA]; Pelham, 2008, J. Res. Lepid. 40: 250; [P&N23]; [CoL] ; [iNaturalist] ; [BOA]
  Larva on Eriogonum nudum, Eriogonum parvifolium var. paynei, E. covilleanum [BNA]
- E. e. langstoni (Shields, 1975) California
  - Shijimiaeoides enoptes langstoni Shields, 1975; Bull. Allyn Mus. 28: 23; TL: California, Mono Co., 1.6 rd mi N of Mono-Inyo Co. line, Hwy 395 ca 6 rd mi S Sherwin Summit, ca. 5700'
  - Euphilotes enoptes langstoni; [NACL], #4367c; Pelham, 2008, J. Res. Lepid. 40: 250; [iNaturalist] ; [BOA]
  - Euphilotes langstoni; [MPG] ; [P&N23]
- E. e. aridorum Austin, 1998 Nevada
  - Euphilotes enoptes aridorum Austin, 1998; Syst. W. N. Am. Butts. (45): 554; TL: Nevada, Washoe Co., Peavine Peak Road at US 395, 1646m
  - Euphilotes enoptes aridorum; Pelham, 2008, J. Res. Lepid. 40: 249; [P&N23]; [iNaturalist] ; [BOA]
- E. e. opacapulla Austin, 1998 Nevada
  - Euphilotes enoptes opacapulla Austin, 1998; Syst. W. N. Am. Butts. (45): 554; TL: Nevada, Clark Co., Newberry Mtns, Grapewine Canyon, 320m
  - Euphilotes dammersi opacapulla; [P&N23]
  - Euphilotes enoptes opacapulla; Pelham, 2008, J. Res. Lepid. 40: 250; [iNaturalist] ; [BOA]
- E. e. primavera Austin, 1998 Nevada
  - Euphilotes enoptes primavera Austin, 1998; Syst. W. N. Am. Butts. (45): 555; TL: Nevada, Mineral Co., Wassuk Range, US Hwy 95, 9.0 road miles S of Schurz, Penrod Canyon, 1463m
  - Euphilotes enoptes primavera; Pelham, 2008, J. Res. Lepid. 40: 250; [P&N23]; [iNaturalist] ; [BOA]
- E. e. arenacola Pratt & Emmel, 1998 California
  - Euphilotes enoptes arenacola Pratt & Emmel, 1998; Syst. W. N. Am. Butts. (16): 210; TL: California, Monterey Co., Marina (sand dunes), 5mi N of Monterey
  - Euphilotes enoptes arenacola; Pelham, 2008, J. Res. Lepid. 40: 250; [P&N23]; [iNaturalist] ; [BOA]
- E. e. cryptorufes Pratt & Emmel, 1998 California
  - Euphilotes enoptes cryptorufes Pratt & Emmel, 1998; Syst. W. N. Am. Butts. (16): 211; TL: California, Riverside Co., S facing slopes of Pyramid Mt, San Jacinto Mtns, 1500-2100m
  - Euphilotes enoptes cryptorufes; [NL4A], #1092a; Pelham, 2008, J. Res. Lepid. 40: 250; [P&N23]; [iNaturalist]
Euphilotes columbiae (Mattoni, 1954) Washington, N.Oregon. See [maps]
- Philotes enoptes columbiae Mattoni, 1954; Bull. Sth. Calif. Acad. Sci. 53 (3): 162; TL: Columbia River nr Brewster, Okanogan Co., Washington
- Euphilotes columbiae; [BNA]; Pelham, 2008, J. Res. Lepid. 40: 250; [MPG] ; [P&N23]; [iNaturalist] ; [BOA] ; [BOA] ; [BAMONA]
- Euphilotes enoptes columbiae; [NACL], #4367g; [CoL]
Larva on Eriogonum compsitum [BNA]
Euphilotes ancilla (Barnes & McDunnough, 1918) SE.Oregon. See [maps]
- Philotes battoides ancilla Barnes & McDunnough, 1918; Contr. nat. Hist. Lepid. N. Am. 4 (2): 79; TL: Eureka, Utah
- Euphilotes ancilla barnesi; [P&N23]; [iNaturalist]
- Euphilotes ancilla campestris; [P&N23]; [iNaturalist]
- Euphilotes ancilla cryptica; [P&N23]; [iNaturalist]
- Euphilotes ancilla montosa; [P&N23]; [iNaturalist]
- Euphilotes ancilla; [Opler]; [BNA]; Pelham, 2008, J. Res. Lepid. 40: 250; [MPG] ; [P&N23]; [iNaturalist] ; [BOA] ; [BOA] ; [BAMONA]
- Euphilotes enoptes ancilla; [NACL], #4367f; [CoL]
Larva on Eriogonum umbellatum, Eriogonum flavum var. crassifolium, E. ovalifolium, E. heracleoides, E. lobii var. robustius, E. strictum var. proliferum [BNA]
- E. a. ancilla Utah
  - Euphilotes ancilla ancilla; Pelham, 2008, J. Res. Lepid. 40: 250; [P&N23]; [iNaturalist] ; [BOA]
- E. a. giulianii Pratt & Emmel, 1998 Nevada
  - Euphilotes ancilla giulianii Pratt & Emmel, 1998; Syst. W. N. Am. Butts. (16): 214; TL: Nevada, Nye Co., Grapevine Mtns, ridge 0.8 km SE of Wahguyhe Peak, 2300m
  - Euphilotes ancilla giulianii; Pelham, 2008, J. Res. Lepid. 40: 250; [P&N23]; [iNaturalist]
- E. a. pseudointermedia Pratt & Emmel, 1998 California
  - Euphilotes ancilla pseudointermedia Pratt & Emmel, 1998; Syst. W. N. Am. Butts. (16): 214; TL: California, Nevada Co., Castle Peak nr Donner Pass, 2700m
  - Euphilotes ancilla pseudointermedia; Pelham, 2008, J. Res. Lepid. 40: 251; [P&N23]; [iNaturalist] ; [BOA]
- E. a. gilvatunica Austin, 1998 Nevada
  - Euphilotes ancilla gilvatunica Austin, 1998; Syst. W. N. Am. Butts. (45): 553; TL: Nevada, Washoe Co., Nevada State Route 341, 3.3 road miles E of US 395, 1525m
  - Euphilotes ancilla gilvatunica; Pelham, 2008, J. Res. Lepid. 40: 251; [P&N23]; [iNaturalist] ; [BOA]
- E. a. shieldsi Austin, 1998 Nevada
  - Euphilotes ancilla shieldsi Austin, 1998; Syst. W. N. Am. Butts. (45): 553; TL: Nevada, Esmeralda Co., White Mtns, Trail Canyon Rd, 9.5 road miles W of Nevada state route 264, 2286m
  - Euphilotes ancilla shieldsi; Pelham, 2008, J. Res. Lepid. 40: 251; [P&N23]; [iNaturalist] ; [BOA]
- E. a. purpura Austin, 1998 Nevada
  - Euphilotes ancilla purpura Austin, 1998; Syst. W. N. Am. Butts. (45): 552; TL: Nevada, Clark Co., Spring Mtns, Willow Creek, 1829m
  - Euphilotes ancilla purpura; Pelham, 2008, J. Res. Lepid. 40: 251; [P&N23]; [iNaturalist] ; [BOA]

RITA-complex

Rita; Pratt & Emmel, 1998, Syst. W. N. Am. Butts. (17): 271; Pelham, 2008, J. Res. Lepid. 40: 251

Euphilotes rita (Barnes & McDunnough, 1916) Nevada, California, Rockies - New Mexico, W.Texas. See [maps] Desert Buckwheat Blue Rita Blue
- Lycaena rita Barnes & McDunnough, 1916; Can. Ent. 48 (7): 223; TL: Arizona
- Euphilotes rita; [NACL], #4369; [BNA]: 405, pl. 37, f. 401; [NAB]; [Opler]; [NL4A], #1093; Pelham, 2008, J. Res. Lepid. 40: 251; [MPG] ; [P&N23]; [CoL] ; [iNaturalist] ; [BOA] ; [BOA] ; [BAMONA]
Larva on Eriogonum effusum, E. flavum, E. leptocladon, E. racemosum, E. wrightii [NAB]
- E. r. rita Arizona, ...
  - Euphilotes rita rita; [NL4A], #1093a; Pelham, 2008, J. Res. Lepid. 40: 251; [P&N23]; [iNaturalist] ; [BOA]
- E. r. coloradensis (Mattoni, [1966]) Colorado
  - Philotes rita coloradensis Mattoni, [1966]; J. Res. Lepid. 4 (2): 88, 90; TL: S of Kendrick, Lincoln Co., Colorado
  - Euphilotes rita coloradensis; [NACL], #4369c; [P&N23]; [iNaturalist] ; [BOA]
Euphilotes pallescens (Tilden & Downey, 1955) NW.Utah, Nevada, California. See [maps] Pale Blue Pallid Blue
- Philotes pallescens Tilden & Downey, 1955; Bull. Southern Calif. Acad. Sci. 54 (1): 25; TL: Little Granite Mts., Dugway Proving Grounds, Tooele Co., Utah
- Euphilotes pallescens; [NACL], #4368; [NAB]; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 251; [MPG] ; [P&N23]; [iNaturalist] ; [BOA] ; [BAMONA]
- Euphilotes rita pallescens; [BNA], 405
- Philotes pallescens; [CoL]
Larva on Eriogonum kearneyi, E. microthecum, E. plumatella [NAB]
- E. p. pallescens
  - Euphilotes pallescens pallescens; Pelham, 2008, J. Res. Lepid. 40: 251; [P&N23]; [iNaturalist] ; [BOA]
- E. p. elvirae (Mattoni, [1966])
  - Philotes rita elvirae Mattoni, [1966]; J. Res. Lepid. 4 (2): 94; TL: 3.5mi SW of Pearblossom, L. A. Co., California
  - Euphilotes pallescens elvirae; [NACL], #4368a; Pelham, 2008, J. Res. Lepid. 40: 252; [P&N23]; [iNaturalist] ; [BOA]
  - Euphilotes rita elvirae; [BNA], 405
- E. p. emmeli (Shields, 1975) Utah
  - Shijimiaeoides rita emmeli Shields, 1975; Bull. Allyn Mus. 28: 16; TL: Utah, Emero Co., summit of rd to 12 mi E of Little Flat Top, 5500'
  - Euphilotes pallescens emmeli; Pelham, 2008, J. Res. Lepid. 40: 251; [P&N23]; [iNaturalist] ; [BOA]
  - Euphilotes rita emmeli; [NACL], #4369b
- E. p. mattonii (Shields, 1975) Nevada
  - Shijimiaeoides rita mattonii Shields, 1975; Bull. Allyn Mus. 28: 20; TL: Nevada, Elko Co., ridge 4 air mi W of Charleston Reservoir, 6600'
  - Euphilotes pallescens mattoni; Pelham, 2008, J. Res. Lepid. 40: 251
  - Euphilotes pallescens mattonii; [P&N23]; [iNaturalist] ; [BOA]
  - Euphilotes rita mattoni; [NACL], #4369a
- E. p. arenamontana Austin, 1998 Nevada
  - Euphilotes pallescens arenamontana Austin, 1998; Syst. W. N. Am. Butts. (45): 556; TL: Nevada, Churchill Co., US Hwy 50, Sand Mountain, 1310m
  - Euphilotes pallescens arenamontana; Pelham, 2008, J. Res. Lepid. 40: 251; [P&N23]; [iNaturalist] ; [BOA]
- E. p. ricei Austin, 1998 Nevada
  - Euphilotes pallescens ricei Austin, 1998; Syst. W. N. Am. Butts. (70): 815
  - Euphilotes pallescens ricei; Pelham, 2008, J. Res. Lepid. 40: 252; [P&N23]; [iNaturalist] ; [BOA]
- E. p. calneva Emmel & Emmel, 1998 California
  - Euphilotes pallescens calneva Emmel & Emmel, 1998; Syst. W. N. Am. Butts. (18): 277; TL: California, Lassen Co., SE Honey Lake Valley
  - Euphilotes pallescens calneva; Pelham, 2008, J. Res. Lepid. 40: 252; [P&N23]; [iNaturalist] ; [BOA]
- E. p. confusa Pratt & Emmel, 1998 California
  - Euphilotes pallescens confusa Pratt & Emmel, 1998; Syst. W. N. Am. Butts. (17): 271; TL: California, Inyo Co., Rest Spring, 6640'
  - Euphilotes pallescens confusa; Pelham, 2008, J. Res. Lepid. 40: 252; [P&N23]; [iNaturalist] ; [BOA]
Euphilotes spaldingi (Barnes & McDunnough, 1917) Nevada, Arizona, Colorado, New Mexico. See [maps]
- Philotes spaldingi Barnes & McDunnough, 1917; Contr. nat. Hist. Lep. N. Am. 3 (4): 216; TL: Provo, Utah
- Euphilotes rita spaldingi; [NACL], #4369d
- Euphilotes spaldingi; [BNA], 405; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 252; [MPG] ; [P&N23]; [CoL] ; [iNaturalist] ; [BOA] ; [BAMONA]
- E. s. spaldingi Utah, ...
  - Euphilotes spaldingi spaldingi; Pelham, 2008, J. Res. Lepid. 40: 252; [P&N23]; [iNaturalist] ; [BOA]
- E. s. pinjuna Scott, 1981 Colorado
  - Euphilotes spaldingi pinjuna Scott, 1981; Papilio (n.s.) 1: 6; TL: Shilling's Spring, Conejos Co., Colorado
  - Euphilotes spaldingi pinjuna; Pelham, 2008, J. Res. Lepid. 40: 252; [P&N23]; [iNaturalist] ; [BOA]

Unmatched external taxa

Euphilotes allyni; [BOA]
Euphilotes ancilla barnesi; [BOA]
Euphilotes baueri vernalis; [BOA]
Euphilotes columbiae columbiae; [BOA]
Euphilotes heracleoides; [MPG] ; [P&N23]; [iNaturalist] ; [BAMONA]
Euphilotes leona; [BOA]
Euphilotes mojave virginensis; [BOA]
Euphilotes oakleyi; [MPG] ; [P&N23]
Euphilotes oakleyi oakleyi; [P&N23]
Euphilotes oakleyi madisonensis; [P&N23]
Euphilotes oakleyi; [iNaturalist]
Euphilotes oakleyi oakleyi; [iNaturalist]
Euphilotes oakleyi madisonensis; [iNaturalist]
Euphilotes speciosa; [BOA]
Euphilotes speciosa bohartorum; [BOA]
Euphilotes speciosa purisima; [BOA]
Euphilotes speciosa septentrionalis; [BOA]
Euphilotes speciosa speciosa; [BOA]
Euphilotes stanfordorum; [MPG] ; [P&N23]; [iNaturalist] ; [BOA] ; [BAMONA]

16.5.2022 (13)

References:

[BAMONA] Opler, Pavulaan, Stanford & Pogue
Butterflies and Moths of North America; Mountain Prairie Information Node
[BOA] Interactive Listing of American Butterflies
Listado Interactivo de la Mariposas Americanas; Butterflies of America
[CEB]
Caucasian Entomological Bulletin; issues (PDF)
[CoL] Catalogue of Life
Bánki, O., Roskov, Y., Döring, M., Ower, G., Hernández Robles, D. R., Plata Corredor, C. A., Stjernegaard Jeppesen, T., Örn, A., Vandepitte, L., Hobern, D., Schalk, P., DeWalt, R. E., Ma, K., Miller, J., Orrell, T., Aalbu, R., Abbott, J., Adlard, R., Aedo, C., et al. (2024). Catalogue of Life Checklist (Version 2024-03-26); https://doi.org/10.48580/dfz8d
[FLMNH] Florida Museum of Natural History
Bulletin of the Allyn Museum
[JRL]
The Journal of Research on the Lepidoptera; JRL Archive
[MPG]
North American Moth Photographers Group
[NHM-BMW] Natural History Museum
Butterflies and Moths of the World; Generic Names and their Type-species
[Opler] Opler & Warren, 2003
Butterflies of North America. 2. Scientific Names List for Butterfly Species of North America, north of Mexico.
[P&N23] Pohl & Nanz, 2023
Annotated Taxonomic Checklist of the Lepidoptera of North America, north of Mexico
[iNaturalist]
iNaturalist
[maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.

Some related literature:

Barnes & McDunnough, 1916
Some new races and species of North American Lepidoptera Can. Ent. 48 (7) : 221-226
Barnes & McDunnough, 1916
Notes on North American diurnal Lepidoptera Contr. nat. Hist. Lep. N. Am. 3 (2) : 49-152, pl. 4-11
Barnes & McDunnough, 1917
Further notes on Philotes battoides and its allies Contr. nat. Hist. Lep. N. Am. 3 (4) : 213-216, pl. 16-17
Barnes & McDunnough, 1918
Notes and new species Contr. nat. Hist. Lepid. N. Am. 4 (2): 61-208, pl. 11-25, (3): 211-278, pl. 26-40
Behr, 1867
Enumeration of the Californian species of Lycaena Proc. Calif. Acad. Sci. 3 (4) : 279-283
Boisduval, 1852
Lépidoptères de la Californie Ann. Soc. ent. Fr. (2) 10 (2) : 275-324
Dyar, 1903
A List of North American Lepidoptera and Key to the Literature of this Order of Insects Bull. U.S. nat. Mus. 52 : 1-723
Edwards, 1871
Descriptions of new species of diurnal Lepidoptera found within the United States Trans. amer. ent. Soc. 3 (3/4) : 205-216
Gunder, 1925
Several New Varieties of and Aberrant Lepidoptera (Rhopalocera) from California Ent. News 36 : 1-9, pl. 1
Gunder, 1927
New transition forms (Lepid., Rhopalocera) Can. Ent. 59 (12) : 281-287
[NACL]; Hodges, 1983
Check List of the Lepidoptera of America North of Mexico Check List Lep. Am. N of Mexico
[NL4A]; Lamas, 2004
Checklist: Part 4A, Hesperioidea - Papilionoidea Atlas of Neotropical Lepidoptera
Langston, 1964
Philotes of central coastal California (Lycaenidae) J. Lep. Soc. 17 (4) : 201-223
[BOW]; Lewis, 1974
Butterflies of the World
Mattoni, [1966]
Distribution and pattern of variation in Philotes rita J. Res. Lepid. 4 (2) : 81-102
Mattoni, [1978]
The Scolitantidini I: two new genera and a generic rearrangement (Lycaenidae) J. Res. Lepid. 16 (4) : 223-242
Mattoni, 1989
The Euphilotes battoides complex: recognition of a species and description of a new subspecies J. Res. Lepid. 27 (3-4) : 173-185
Pelham, 2008
A catalogue of the butterflies of the United States and Canada J. Res. Lepid. 40 : 1-652
Pratt & Emmel, 1998
Revision of the Euphilotes enoptes and E. battoides complexes Syst. W. N. Am. Butts. (16) : 207-270
Pratt & Emmel, 1998
A new subspecies of Euphilotes pallescens (Lepidoptera: Lycaenidae) from the Death Valley region of California Syst. W. N. Am. Butts. (17) : 271-276
[NAB]; Pyle, 1981
Field Guide to North American Butterflies, National Audubon Society
Scott, 1981
New Papilionoidea and Hesperioidea from North America Papilio (n.s.) 1 : 1-12
[BNA]; Scott, 1986
The Butterflies of North America Butts. N. America : 1-583
Scott, 1998
New western North American butterflies Papilio (n.s.) 11 : 1-12
Shields, 1975
Studies on North American Philotes. IV. Taxonomic and biological notes, and new subspecies Bull. Allyn Mus. 28
Stradomsky, 2016
A molecular phylogeny of the subfamily Polyommatinae (Lepidoptera: Lycaenidae) Caucasian Ent. Bull. 12 (1) : 145-156
Tilden & Downey, 1955
A New Species of Philotes from Utah Bull. Southern Calif. Acad. Sci. 54 (1): 25-29
Ugelvig, Vila, Pierce & Nash, 2011
A phylogenetic revision of the Glaucopsyche section (Lepidoptera: Lycaenidae), with special focus on the Phengaris-Maculinea clade Molec. Phyl. Evol. 61 : 237-243
Watson & Comstock, 1920
Notes on American Lepidoptera with descriptions of new varieties Bull. Amer. Mus. 42 (10) : 449-457

If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.

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