Lethe dura (Marshall, 1882) Himalays, W.China, Formosa. See [maps]

• Zophoessa dura Marshall, 1882; J. Asiat. Soc. Bengal 51 Pt.II (2-3): 38, pl. 4, f. 2; TL: Upper Tenasserim

• Lethe (Zophoessa) dura; Lang, 2020, Atalanta 51 (3/4): 333 (list)
• Lethe dura; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 314; [BOW]: pl. 201, f. 13; [BOR], 412; [MRS], 322; Lang, 2020, Atalanta 51 (1/2): 63; [LepIndex] ; [CoL] ; [iNaturalist]

• L. d. dura Yunnan, N.Vietnam, Laos, E.Myanmar, N.Thailand

  • Lethe dura dura; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 314; [BOR], 412; Lang, 2020, Atalanta 51 (1/2): 63; [iNaturalist] ; [Yutaka Inayoshi]
  • Zophoessa dura dura; Huang, 2003, Neue Ent. Nachr. 55: 91 (note)
• L. d. moupiniensis (Poujade, 1884) China (SE.Gansu, S.Shaanxi, Hubei, Zhejiang, Fujian, Jiangxi, Guanglong, Sichuan, Chongqing, Guizhou)

  • Debis moupiniensis Poujade, 1884; Bull. Soc. Ent. Fr. (6) 4: cxl
  • Zophoessa libitina Leech, 1891; Entomologist 24 (Suppl.): 2; TL: Chia-Ting-Fu; Chang-Yang
    • = Lethe dura moupinensis; Lang, 2014, Atalanta 45 (1-4): 172; Lang, 2020, Atalanta 51 (1/2): 64

  • Lethe dura moupinensis[sic, moupiniensis]; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 314; [MRS], 322; [BOR], 412; Lang, 2014, Atalanta 45 (1-4): 172; Lang, 2020, Atalanta 51 (1/2): 64
  • Zophoessa moupinensis[sic, moupiniensis]; Seitz, 1907, Gross-Schmett. Erde 1: 86, pl. 32 a
• L. d. gammiei (Moore, [1892]) NE.India, Bhutan, W.Yunnan, N.Myanmar

  • dura gammiei (Moore, [1892]) (Zophoessa); Lepidoptera Indica 1: 294, pl. 91, f. 3; TL: Bhutan

  • Lethe dura gammiei; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 314; [BOR], 412; Lang, 2020, Atalanta 51 (1/2): 63
• L. d. neoclides Fruhstorfer, 1909 Taiwan

  • Lethe dura neoclides Fruhstorfer, 1909; Soc. ent. 24 (16): 121; TL: Formosa

  • Lethe dura neoclides; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 314; [BOR], 412; [MRS], 322; Lang, 2020, Atalanta 51 (1/2): 64; [iNaturalist]
• L. d. mansonia Fruhstorfer, 1911 NE.Vietnam, Hainan

  • Lethe dura mansonia Fruhstorfer, 1911; Gross-Schmett. Erde 9: 314, pl. 99 a; TL: Manson Mts., N.Vietnam

  • Lethe dura mansonia; [BOR], 412; Lang, 2020, Atalanta 51 (1/2): 63; [Yutaka Inayoshi]

Lethe dataensis (Semper, 1887) Philippines (Luzon, Mindoro). See [maps]

• Debis ? dataensis Semper, 1887; Reisen Philipp. (2): 57, pl. 11, f. 4; TL: NW.Luzon

• Lethe (Zophoessa) dataensis; Lang, 2020, Atalanta 51 (3/4): 333 (list)
• Lethe dataensis; [BOR], 415
• Lethe dura daatensis[sic, dataensis]; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 315

• L. d. halcona (Yamamoto & Takei, 1993) Philippines (Mindanao)

  • Zophoessa dataensis halcona Yamamoto & Takei, 1993; Tyô to Ga 43 84): 255, f. 1-2; TL: Mt. Halcon (1500m), N.Mindanao

Lethe goalpara (Moore, [1866]) Simla Hills, Sikkim, Assam. See [maps] Large Goldenfork

• Zophoessa goalpara Moore, [1866]; Proc. zool. Soc. Lond. 1865 (3): 768

• Lethe (Zophoessa) goalpara; Lang, 2020, Atalanta 51 (3/4): 333 (list)
• Lethe goalpara; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 314, pl. 99 a; [BOW]: pl. 167, f. 5; [BOR], 414; [LepIndex] ; [CoL] ; [iNaturalist]

600x400 (~49Kb) upperside India: Arunachal Pradesh, Taley Valley WLS, 11.11.2005, Photo © Kedar Tokekar

600x400 (~82Kb) underside India: Arunachal Pradesh, Taley Valley WLS, 11.11.2005, Photo © Kedar Tokekar

• L. g. goalpara N.India - Assam

  • ?Lethe goalpara narkunda Fruhstorfer, 1911; Gross-Schmett. Erde 9: 314; TL: Simla

  • Lethe goalpara goalpara; [BOR], 414; [Yutaka Inayoshi]
• L. g. kabruensis Tytler, 1939 Manipur (Kabru)

  • goalpara kabruensis Tytler, 1939; J. Bombay nat. Hist. Soc. 41 (2): 246; TL: Burma

  • Lethe goalpara kabruensis; [BOR], 414; [LepIndex]; [CoL]
• L. g. gana Talbot, 1947 Upper Burma

  • goalpara gana Talbot, 1947; Fauna Br. Ind., Butt. (2nd ed.) 2: 186; TL: Sikkim

  • Lethe goalpara gana; [BOR], 414; [LepIndex]; [CoL]

Lethe yantra Fruhstorfer, 1914 W.China. See [maps]

• Lethe yantra Fruhstorfer, 1914; Ent. Rundschau 31 (5): 25; TL: Yunnan

• Lethe (Zophoessa) yantra; Lang, 2020, Atalanta 51 (3/4): 333 (list)
• Lethe yantra; [MRS], 334; [LepIndex] ; [CoL] ; [iNaturalist]

• L. y. distincta Mell, 1923

  • Lethe callipteris distincta Mell, 1923; Dt. ent. Zs. 1923 (2): 154

  • Lethe yantra distincta; [MRS], 334

Lethe moelleri (Elwes, 1887) Bhutan, Sikkim - Guanxi. See [maps] Moeller's Silverfork

• Zophoessa mölleri [= moelleri] Elwes, 1887; Proc. zool. Soc. Lond. 1887: 445

• Lethe (Zophoessa) moelleri; Lang, 2020, Atalanta 51 (3/4): 333 (list)
• Lethe moelleri; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 314, pl. 97 d; [BOR], 416; [MRS], 328; [LepIndex] ; [CoL]

• L. m. bruno Tytler, 1939 NE.Burma

  • Lethe moelleri bruno Tytler, 1939; J. Bombay nat. Hist. Soc. 41 (2): 246; TL: Hthawgaw, NE Burma

  • Lethe moelleri bruno; Huang, 2001, Neue Ent. Nachr. 51: 95 (note); [LepIndex]; [CoL]

Lethe gregoryi Watkins, 1927 Yunnan. See [maps]

• Lethe (Magula) helle gregoryi Watkins, 1927; Ann. Mag. nat. Hist. (9) 19: 323; TL: Yangtze Valley, E of Janula, Yunnan
• Lethe moelleri bitaensis Yoshino, 1999; Neo Lepidoptera 4: 7, f. 49-50; TL: Bita Lake, Zhongdian, NW.Yunnan
  • = ?Lethe procne bitaensis Yoshino, 1999; Huang, 2001, Neue Ent. Nachr. 51: 95
  • = ; Lang, 2015, Zootaxa 4048 (4): 575; Lang, 2016, Atalanta 47 (3/4): 343

• Lethe (Zophoessa) gregoryi; Lang, 2020, Atalanta 51 (3/4): 333 (list)
• Lethe bitaensis; Yoshino, 2003, Futao (43): 8
• Lethe gregoryi; Lang, 2015, Zootaxa 4048 (4): 575; Lang, 2016, Atalanta 47 (3/4): 343; [iNaturalist]
• Lethe helle gregoryi; [MRS], 334

• L. g. gesangdawai Huang, 2001 SE.Tibet (Nujiang)

  • Lethe gesangdawai Huang, 2001; Neue Ent. Nachr. 51: 95, f. 86, pl. 7, f. 55; TL: 10km E of Nuola, Nujiang Valley, SE.Tibet
    • = Lethe bitaensis; Yoshino, 2003, Futao (43): 8

  • Lethe gregoryi gesangdawai; Lang, 2015, Zootaxa 4048 (4): 575

Lethe procne (Leech, 1891) -. See [maps]

• Zophoessa procne Leech, 1891; Entomologist 24 (Suppl.): 2; TL: Wa-Shan; Huang-Mu-Chang; Ta-Tsien-Lu

• Lethe (Zophoessa) procne; Lang, 2020, Atalanta 51 (3/4): 333 (list)
• Lethe procne; Lang & Liu, 2014, Atalanta 45 (1-4): 168; [LepIndex] ; [CoL] ; [iNaturalist]
• Zophoessa procne; Seitz, 1907, Gross-Schmett. Erde 1: 87, pl. 32 b; Huang, Wu & Yuan, 2003, Neue Ent. Nachr. 55: 151 (note)

• L. p. procne China (N.Yunnan, Sichuan, S.Shaanxi)

  • Lethe procne procne; Lang, 2016, Atalanta 47 (1/2): 230
• L. p. abaensis Lang, 2016 China (N.Sichuan, S.Gansu)

  • Lethe procne abaensis Lang, 2016; Atalanta 47 (1/2): 230; TL: China, Sichuan, Maoxian, Songping-gou, 2400-2900m
• L. p. shunana Lang, 2016 China (SW.Sichuan, N.Yunnan)

  • Lethe procne shunana Lang, 2016; Atalanta 47 (1/2): 231; TL: China, Sichuan, Yanbian, Gesala, 3000m

Lethe paraprocne Lang & Liu, 2014 -. See [maps]

• Lethe paraprocne Lang & Liu, 2014; Atalanta 45 (1-4): 167; TL: Sichuan, Heizhugou, Ebian

• Lethe (Zophoessa) paraprocne; Lang, 2020, Atalanta 51 (3/4): 333 (list)

• L. p. paraprocne Sichuan

  • Lethe paraprocne paraprocne; Lang, 2016, Atalanta 47 (1/2): 231; Huang, 2019, Neue Ent. Nachr. 78: 208
• L. p. kawagarboensis Lang & Liu, 2014 NW.Yunnan

  • Lethe paraprocne kawagarboensis Lang & Liu, 2014; Atalanta 45 (1-4): 167; TL: Yunnan, Deqin, Mingyong village, 2500m

  • Lethe paraprocne kawagarboensis; Lang, 2016, Atalanta 47 (1/2): 231; Huang, 2019, Neue Ent. Nachr. 78: 208
• L. p. shunana Lang, 2016 S.Sichuan

  • Lethe procne shunana Lang, 2016; ; TL: Gesala

  • Lethe paraprocne shunana; Huang, 2019, Neue Ent. Nachr. 78: 208
• L. p. zhongdiana Huang, 2019 NW.Yunnan

  • Lethe paraprocne zhongdiana Huang, 2019; Neue Ent. Nachr. 78: 208; TL: China, Yunnan, Shangri-la City, ca. 3150m

Lethe ocellata (Poujade, 1885) Sichuan, Tibet, Sikkim, Manipur. See [maps] Dismal Mystic

• Debis ocellata Poujade, 1885; Bull. Soc. Ent. Fr. (6) 5: x; TL: Mou-pin, Sichuan
• Lethe simulans Leech, 1891; Entomologist 24 (Suppl.): 23; TL: Omei-Shan
  • = ; Seitz, 1907, Gross-Schmett. Erde 1: 85; Huang, 2014, Atalanta 45 (1-4): 152

• Lethe (Zophoessa) ocellata; Lang, 2020, Atalanta 51 (3/4): 333 (list)
• Lethe ocellata; Leech, [1892], Butts China Japan Corea (1): 34-35; Seitz, 1907, Gross-Schmett. Erde 1: 85, pl. 31 c; Seitz, [1909], Gross-Schmett. Erde 1: 85, pl. 31 c; Draeseke, 1925, Dt. ent. Z. Iris 39: 52; Lee & Zhu, 1992, Atlas of Chinesee butteflies: 93-94; [MRS], 333; Huang, 2014, Atalanta 45 (1-4): 152; [LepIndex] ; [CoL] ; [iNaturalist] ; [Yutaka Inayoshi]
• Zophoessa ocellata; de Lesse, 1956, Ann. Soc. ent. Fr. 125: 78; Huang, 2002, Atalanta 33 (3/4): 365, pl. 21, f. 4, 8; Huang, 2003, Neue Ent. Nachr. 55: 91 (note), f. 140; Huang, Wu & Yuan, 2003, Neue Ent. Nachr. 55: 145, pl. 12, f. 1

• L. o. mon Yoshino, 2008 N.Vietnam

  • Lethe ocellata mon Yoshino, 2008; Futao (54): 10, pl. 2, f. 11-12; TL: Sapa, 2000m, N.Vietnam
    • = Lethe ocellata; Huang, 2014, Atalanta 45 (1-4): 152

Lethe nigrifascia Leech, 1890 China (Hubei, Hunan, Henan, Shaanxi, Gansu, Sichuan). See [maps]

• Lethe nigrifascia Leech, 1890; Entomologist 23: 28; TL: Hubei, Chang-yang

• Lethe (Zophoessa) nigrifascia; Lang, 2020, Atalanta 51 (3/4): 333 (list)
• Lethe nigrifascia; Leech, [1892], Butts China Japan Corea (1): 33; Seitz, 1907, Gross-Schmett. Erde 1: 85, pl. 31 e; Huang, 1999, Lambillionea 99 (1): 129-131, f. 1b, 2b, 3b; Huang, 1998, Neue Ent. Nachr. 41: 225 (note), pl. 4, f. 3c, 4c; [BOW]: pl. 202, f. 6; [MRS], 334; Lang & Wang, 2016, Atalanta 47 (1/2): 225 (note); [LepIndex] ; [CoL] ; [iNaturalist]
• Zophoessa nigrifascia; de Lesse, 1956, Ann. Soc. ent. Fr. 125: 79; Huang, 2003, Neue Ent. Nachr. 55: 91 (note), f. 138; Huang, Wu & Yuan, 2003, Neue Ent. Nachr. 55: 148, pl. 12, f. 2

• L. n. ebiana Lang, 2014 Sichuan

  • Lethe nigrifascia ebiana Lang, 2014; Neue Ent. Nachr. 70: 299; TL: China, Sichuan, Ebian, Heizhugou, 1800-2000m
  • Lethe nigrifascia ab. fasciata Seitz, 1907; Gross-Schmett. Erde 1: 85, pl. 31 d; TL: Sichuan, Pu-tsu-fong
    • = ; Lang, 2014, Neue Ent. Nachr. 70: 299
    • = ?Lethe liyufeii; Huang, 2014, Atalanta 45 (1-4): 151

  • Lethe nigrifascia ebiana; Lang & Wang, 2016, Atalanta 47 (1/2): 225 (note)
• L. n. lojishana Lang, 2016 Sichuan

  • Lethe nigrifascia lojishana Lang, 2016; Atalanta 47 (3/4): 341; TL: China, Sichuan, Puge, Mt Luojishan, 2600m

Lethe jalaurida (de Nicéville, 1881) W.China, Kulu - Sikkim. See [maps] Small Silverfork

• Zophoessa jalaurida de Nicéville, 1881; J. Asiat. Soc. Bengal 49 Pt.II (4): 245; TL: Jalauri Pass, NW.Himalayas

• Lethe (Zophoessa) jalaurida; Lang, 2020, Atalanta 51 (3/4): 333 (list)
• Lethe jalaurida; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 313; [BOW]: pl. 167, f. 6; [BOR], 416; [MRS], 328; Das, Lang, Singh & Chandra, 2020, Zootaxa 4789 (1): 282; [LepIndex] ; [CoL] ; [iNaturalist]
• Zophoessa jalaurida; Seitz, 1907, Gross-Schmett. Erde 1: 87, pl. 32 c

• L. j. jalaurida W.Himalaya (foothills)

  • Lehte jalaurida jalaurida; Huang, 2001, Neue Ent. Nachr. 51: 94 (note)
  • Lethe jalaurida jalaurida; [BOR], 416; [iNaturalist]

Lethe gelduba Fruhstorfer, 1911 -. See [maps]

• Lethe jalaurida gelduba Fruhstorfer, 1911; Gross-Schmett. Erde 9: 313 (not preocc. Lethe sidonis f. gelduba Fruhstorfer, 1911)
• Lethe jalaurida plistia Fruhstorfer, 1911; Gross-Schmett. Erde 9: 394 (repl. Lethe jalaurida gelduba Fruhstorfer, 1911)
  • = ; Das, Lang, Singh & Chandra, 2020, Zootaxa 4789 (1): 285

• Lethe (Zophoessa) gelduba; Lang, 2020, Atalanta 51 (3/4): 333 (list)
• Lethe gelduba; Das, Lang, Singh & Chandra, 2020, Zootaxa 4789 (1): 285
• Lethe jalaurida gelduba; [BOR], 416; Huang, 2001, Neue Ent. Nachr. 51: 94 (note); [iNaturalist]
• Lethe plistia; Lang, 2014, Neue Ent. Nachr. 70: (299-308)

• L. g. gelduba W.Sichuan, N.Yunnan

  • Lethe gelduba gelduba; Das, Lang, Singh & Chandra, 2020, Zootaxa 4789 (1): 285
• L. g. nuolaensis Huang, 2001 SE.Tibet

  • Lethe jalaurida nuolaensis Huang, 2001; Neue Ent. Nachr. 51: 94, pl. 7, f. 52; TL: 5km E of Nuola, Nujiang Valley, SE.Tibet

  • Lethe gelduba nuolaensis; Das, Lang, Singh & Chandra, 2020, Zootaxa 4789 (1): 285
  • Lethe jalaurida nuolaensis; [iNaturalist]

Lethe maitrya de Nicéville, 1881 -. See [maps] Barred Woodbrown

• Lethe maitrya de Nicéville, 1881; J. Asiat. Soc. Bengal 49 Pt.II (4): 245; TL: Jalauri pass, NW.Himalayas

• Lethe (Zophoessa) maitrya; Lang, 2020, Atalanta 51 (3/4): 333 (list)
• Lethe maitrya; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 313, pl. 97 c; [BOW]: pl. 167, f. 15; [BOR], 414; [MRS], 335; [LepIndex] ; [CoL] ; [iNaturalist]

• L. m. maitrya W.Himalayas, Nepal, Sikkim, Bhutan

  • Lethe maitrya maitrya; Huang, 2001, Neue Ent. Nachr. 51: 96 (note)
• L. m. thawgawa Tytler, 1939 NE.Burma, Nujiang Valley

  • Lethe maitrya thawgawa Tytler, 1939; J. Bombay nat. Hist. Soc. 41 (2): 245; TL: Burma

  • Lethe maitrya thawgawa; Huang, 2001, Neue Ent. Nachr. 51: 96 (note), pl. 7, f. 56; [LepIndex]; [CoL]
  • Zophoessa maitrya thawgawa; Huang, 2003, Neue Ent. Nachr. 55: 91 (note)

  Misidentified/misapplied:

  • Lethe maitrya thawgawa; D'Abrera, 1993, 124 (Tsekou)
    • = Lethe maitrya lijiangensis; Huang, 2001, Neue Ent. Nachr. 51: 96
  • Lethe maitrya thawgawa; Huang, 1998, Neue Ent. Nachr. 41: 251, pl. 4, f. 1d, 2d
    • = Lethe maitrya lijiangensis; Huang, 2001, Neue Ent. Nachr. 51: 96
• L. m. metokana Huang, 1998 SE.Tibet (Metok)

  • Lethe maitrya metokana Huang, 1998; Neue Ent. Nachr. 41: 223, pl. 4, f. 1a-c, 2a-c; TL: Metok
• L. m. lijiangensis Huang, 2001 NW.Yunnan

  • Lethe maitrya lijiangensis Huang, 2001; Neue Ent. Nachr. 51: 96; TL: Lijiang, NW.Yunnan, 3500m

Lethe siderea Marshall, 1881 Sikkim - W.China. See [maps] Scarce Woodbrown

• Lethe siderea Marshall, 1881; J. Asiat. Soc. Bengal 49 Pt.II (4): 246; TL: Sikkim

• Lethe (Zophoessa) siderea; Lang, 2020, Atalanta 51 (3/4): 333 (list)
• Lethe siderae; [LepIndex] ; [CoL]
• Lethe siderea; Seitz, 1907, Gross-Schmett. Erde 1: 85; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 312; [BOR], 415; [MRS], 343; [iNaturalist]
• Zophoessa siderea; ; [iNaturalist]

• L. s. siderea

  • Lethe siderea siderea; [Yutaka Inayoshi]
• L. s. kanoi Esaki & Nomura, 1937 Taiwan

  • Lethe siderea kanoi Esaki & Nomura, 1937; Zephyrus 7: 107; TL: Formosa

  • Lethe siderea kanoi; [MRS], 343

Larva on Arundinaria nitakayamensis [MRS]

Lethe callipteris (Butler, 1877) Japan. See [maps]

• Neope callipteris Butler, 1877; Ann. Mag. nat. Hist. (4) 19 (109): 92
• ?suffusa Esaki, 1924²; Insec. Inscit. Mens. 12: 56

• Harima callipteris; Kudrna, 1974, Atalanta 5: 101; [BRU], 184
• Lethe (Zophoessa) callipteris; Lang, 2020, Atalanta 51 (3/4): 333 (list)
• Lethe callipteris; Seitz, 1907, Gross-Schmett. Erde 1: 86, pl. 31 e; [LepIndex] ; [CoL] ; [iNaturalist]
• Zophoessa callipteris; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 44

720x670 (~52Kb) The Kojoro Lake (Kojoroga-ike) bank on Mt. Hira, 1060 m altitude, Katsuragawa Sakashita-cho, Ohtsu Cty, Shiga Prefecture, Honshu, Japan. 2nd August 2002, Photo © Oleg Kosterin

636x636 (~58Kb) underside The Kojoro Lake (Kojoroga-ike) bank on Mt. Hira, 1060 m altitude, Katsuragawa Sakashita-cho, Ohtsu Cty, Shiga Prefecture, Honshu, Japan. 2nd August 2002, Photo © Oleg Kosterin

Larva on Sasa [BRU], Sasa kurilensis [RFEL]

• L. c. diluta Esaki, 1924 Japan, ...

  • callipteris diluta Esaki, 1924; Insec. Inscit. Mens. 12: 56, pl. 3, f. 3-4; TL: Hokkaido

  • Zophoessa callipteris diluta; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 44
• L. c. karafutonis Matsumura, 1925 Sakhalin

  • Lethe callipteris karafutonis Matsumura, 1925; J. Coll. Agric. Hokkaido imp. Univ. 15: 93, pl. 9, f. 13 ♀; TL: Saghalin
  • minima Esaki, 1924; Insec. Inscit. Mens. 12: 56, pl. 3, f. 8; TL: Sakhalin
  • ?minima Esaki & Nakahara, 1924
• L. c. obscura Nakahara, 1926 Kuriles

  • Lethe callipteris obscura Nakahara, 1926; Ins. Inscit. Mens. 14: 50; TL: Kuriles

Lethe europa (Fabricius, 1775) S.India, Dun - Assam, Burma, N.India, Japan, Hong Kong. See [maps] Bamboo Tree-brown

• Papilio europa Fabricius, 1775; Syst. Ent.: 500, no. 247; TL: S.India

• Lethe (Lethe) europa; Lang, 2020, Atalanta 51 (3/4): 334 (list)
• Lethe europa; Wood-Mason & de Nicéville, 1881, J. Asiat. Soc. Bengal 49 Pt.II (4): 226; Seitz, 1907, Gross-Schmett. Erde 1: 84, pl. 30 d e; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 315; [BOW]: pl. 167, f. 3; [BOR], 416; [MRS], 327; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 176; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]

512x341 (~27Kb) underside India: Guwahati, 16.11.2005, Photo © Kedar Tokekar

Larva on Bambusa multiplex, Microstegium ciliatum, Leleba multiplex [MRS], Bambusa , Dinochloa , Microstegium , Phyllostachys , Pleioblastus Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 176

• L. e. europa Java, Bawean, Kangean, Bali

  • Lethe europa europa; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 316, pl. 99 d; [BOR], 416; [iNaturalist]
• L. e. beroe (Cramer, [1775]) S.China, ?Hong Kong

  • Papilio beroe Cramer, [1775]; Uitl. Kapellen 1 (1-7): pl. 79, f. C, D
    • = Lethe europa; Seitz, 1907, Gross-Schmett. Erde 1: 84

  • Lethe europa beroe; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 316; [BOR], 417; [MRS], 327; [iNaturalist]
• L. e. arcuata Butler, 1868 N.Sulawesi, Lutungan I.

  • Lethe arcuata Butler, 1868; Cat. diurn. Lep. Satyridae Brit. Mus.: 114, pl. 2, f. 3; TL: Celebes

  • Lethe arete arcuata; [BOR], 417
  • Lethe europa arcuata; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 176
• L. e. tamuna de Nicéville, 1887 Nicobars

  • Lethe tamuna de Nicéville, 1887; Proc. zool. Soc. Lond. 1887 (3): 449, pl. 39, f. 6 ♀; TL: Little Nicobar

  • Lethe europa tamuna; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 316; [BOR], 416
• L. e. pavida Fruhstorfer, 1908 Japan, Taiwan

  • Lethe europa pavida Fruhstorfer, 1908; Ent. Zs. 22 (35): 141; TL: Kagi

  • Lethe europa pavida; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 316; [BOR], 417; [MRS], 327; [iNaturalist]

  Larva on Phyllostachys makinoi, Bambusa multiplex, B. oldhamii [MRS]

• L. e. niladana Fruhstorfer, 1911 N.India - Burma, Thailand

  • Lethe europa niladana Fruhstorfer, 1911; Gross-Schmett. Erde 9: 315; TL: "Kuamon - Burma"

  • Lethe europa niladana; [BOR], 416; [MRS], 327; [iNaturalist] ; [Yutaka Inayoshi]
• L. e. nagaraja Fruhstorfer, 1911 S.Sulawesi, Selajar

  • Lethe arete nagaraja Fruhstorfer, 1911; Gross-Schmett. Erde 9: 316

  • Lethe arete nagaraja; [BOR], 417
  • Lethe europa nagaraja; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 176
• L. e. anatha Fruhstorfer, 1911 Sula Is.

  • Lethe arete anatha Fruhstorfer, 1911; Gross-Schmett. Erde 9: 316; TL: Sula I.

  • Lethe arete anatha; [BOR], 417
  • Lethe europa anatha; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 176
• L. e. advipa Fruhstorfer, 1911 Bachan, Halmahera, Ternate

  • Lethe arete advipa Fruhstorfer, 1911; Gross-Schmett. Erde 9: 316

  • Lethe arete advipa; [BAUR], 277
• L. e. velitra Fruhstorfer, 1911 Sanghie

  • Lethe arete velitra Fruhstorfer, 1911; Gross-Schmett. Erde 9: 316; TL: Sangir

  • Lethe arete velitra; [BOR], 417
  • Lether europa velitra; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 176
• L. e. gada Fruhstorfer, 1911 S.Yunnan

  • Lethe europa f. gada Fruhstorfer, 1911; Gross-Schmett. Erde 9: 316; TL: Siam; Tonkin

  • Lethe europa gada; Huang & Xue, 2004, Neue Ent. Nachr. 57: 140 (name)
• L. e. ragalva Fruhstorfer, 1911 S.India

  • Lethe europa ragalva Fruhstorfer, 1911; Gross-Schmett. Erde 9: 316; TL: Karwar

  • Lethe europa ragalva; [BOR], 416
• L. e. nudgara Fruhstorfer, 1911 Andamans

  • Lethe europa nudgara Fruhstorfer, 1911; Gross-Schmett. Erde 9: 316, pl. 96 c; TL: Andamans

  • Lethe europa nudgara; [BOR], 416; [iNaturalist]
• L. e. kumara Fruhstorfer, 1911 Enggano

  • Lethe europa kumara Fruhstorfer, 1911; Gross-Schmett. Erde 9: 316; TL: Engano

  • Lethe europa kumara; [BOR], 416
• L. e. mahamaya Fruhstorfer, 1911 Lombok

  • Lethe europa mahamaya Fruhstorfer, 1911; Gross-Schmett. Erde 9: 316, pl. 96 d; TL: Lombok

  • Lethe europa mahamaya; [BOR], 417
• L. e. alaca Fruhstorfer, 1911 Palawan

  • Lethe europa alaca Fruhstorfer, 1911; Gross-Schmett. Erde 9: 316

  • Lethe europa alaca; [BOR], 417
• L. e. cevanna Fruhstorfer, 1911 Philippines (Mindanao)

  • Lethe europa cevanna Fruhstorfer, 1911; Gross-Schmett. Erde 9: 316; TL: Mindanao

  • Lethe europa cevanna; [BOR], 417
• L. e. malaya Corbet, 1941 Peninsular Malaya, Singapore

  • Lethe europa malaya Corbet, 1941; Entomologist 74 (6): 133; TL: Malaysia

  • Lethe europa malaya; [BOR], 416; [BMP]: 124, pl. 15, f. 7-9; [LepIndex]; [CoL] ; [iNaturalist] ; [Yutaka Inayoshi]

  1211x1031 (~343Kb) ♀ underside Thailand, Phuket, a garden in the Nai Yang beach area. 16th February 2009, Photo © Oleg Kosterin

Lethe rohria (Fabricius, 1787) Afghanistan, Ceylon, S.India - Pachmarhi, Kashmir - Assam, Burma. See [maps] Common Treebrown

• Papilio rohria Fabricius, 1787; Mantissa Insectorum 2: 45; TL: S.India
• Lethe rohria f. stenopoides Mell, 1942; Arch. Naturgesch. (N.F.) 11: 272

• Lethe (Lethe) rohria; Lang, 2020, Atalanta 51 (3/4): 334 (list)
• Lethe rohria; Seitz, 1907, Gross-Schmett. Erde 1: 84, pl. 30 e; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 315; Moore, 1878, Proc. zool. Soc. Lond. 1878 (4): 824; [BOW]: pl. 167, f. 13; [BOR], 418; [MRS], 327; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]

600x400 (~55Kb) underside India: Pune, 18.12.2005 © Kedar Tokekar

• L. r. rohria Burma

  • Debis dyrta C. & R. Felder, [1867]; Reise Fregatte Novara, Bd 2 (Abth. 2) (3): 497; TL: Bengalia
  • ?dyrta Bingham, 1905²; Fauna Brit. Ind., Butts (ed. 1) 1: 80

  • Lethe dyrta; Seitz, 1907, Gross-Schmett. Erde 1: 84, pl. 30 d; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 316; [LepIndex] ; [CoL]
  • Lethe rohria rohria; [BOR], 418; [iNaturalist] ; [Yutaka Inayoshi]
  • rohria r. dyrta; [BIR], (Kashmir - Kumaon)
  • rohria r. rohria; [BIR], (Sikkim - Burma)

  945x938 (~189Kb) ♀ underside Cambodia, Mondulkiri Province, Dak Dam village environs, 16th June 2014, Photo © Oleg Kosterin

• L. r. neelgheriensis (Guérin-Méneville, 1843) Ceylon, S.India - S.Bihar, Jubbulpore, Pachmarhi, Konkan

  • Satyrus neelgheriensis Guérin-Méneville, 1843; in Delessert, Souvenirs Voy. Inde (2): 74, pl. 21, f. 1, 1a

  • Debis neelgheriensis; Hewitson, 1863, Ill. exot. Butts [4] (Debis II-III): [35], pl. [19], f. 6
  • Lethe dyrta neelgheriensis; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 317
  • Lethe neelgherriensis[sic, neelgheriensis]; Moore, [1881], Lepid. Ceylon 1 (1): 16, pl. 7, f. 1, 1a
  • Lethe rohria neelgheriensis; [BOR], 418; [iNaturalist]
  • rohria r. nilgiriensis[sic, neelgheriensis]; [BIR]
• L. r. yoga Fruhstorfer, 1911 Ceylon

  • lethe dyrta yoga Fruhstorfer, 1911; Gross-Schmett. Erde 9: 317; TL: Ceylon

  • Lethe rohria yoga; [BOR], 418
• L. r. permagnis Fruhstorfer, 1911 S.China

  • Lethe dyrta permagnis Fruhstorfer, 1911; Gross-Schmett. Erde 9: 317

  • Lethe rohria permagnis; [BOR], 418; [MRS], 327
• L. r. anunda Fruhstorfer, 1911 Java, Bali

  • Lethe dyrta anunda Fruhstorfer, 1911; Gross-Schmett. Erde 9: 317; TL: Bali, Java

  • Lethe rohria anunda; [BOR], 418
• L. r. daemoniaca Fruhstorfer, 1908 Taiwan

  • Lethe dyrta daemoniaca Fruhstorfer, 1908; Ent. Zs. 22 (35): 141

  • Lethe dyrta daemoniaca; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 317
  • Lethe rohria daemoniaca; [BOR], 418; [MRS], 327

  Larva on Miscanthus sinesis, M. flavidus, Brachiaria mutica, Imperata cylindrica [MRS]

• L. r. sambaluna Fruhstorfer, 1911 Lombok

  • Lethe dyrta sambaluna Fruhstorfer, 1911; Gross-Schmett. Erde 9: 317, pl. 97 b; TL: Lombok

  • Lethe rohria sambaluna; [BOR], 418

Lethe darena (C. & R. Felder, [1867]) Java, Sumatra, Borneo. See [maps]

• Debis darena C. & R. Felder, [1867]; Reise Fregatte Novara, Bd 2 (Abth. 2) (3): 498, pl. 68, f. 4-5; TL: Java

• Lethe (Lethe) darena; Lang, 2020, Atalanta 51 (3/4): 334 (list)
• Lethe darena; de Nicéville, 1893, J. Bombay Nat. Hist. Soc. 8 (1): 40, pl. K, f. 7; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 322; [BOW]: pl. 167, f. 1; [BOR], 424; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]

• L. d. darena W.Java (Gunong Gede)

  • Lethe darena darena; [BOR], 424
• L. d. sumatrensis Staudinger, [1897] Sumatra (Battak Mts)

  • Lethe darena var. sumatrensis Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2): 228

  • Lethe darena sumatrensis; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 322, pl. 98 d; [BOR], 424
• L. d. borneensis Staudinger, [1897] Borneo

  • Lethe darena var. borneensis Staudinger, [1897]; Dt. Ent. Z. Iris 9 (2): 228; TL: Kina Balu

  • Lethe darena borneensis; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 322, pl. 98 e; [BOR], 424; [iNaturalist]

Lethe verma (Kollar, [1844]) Kashmir - Assam, Burma. See [maps] Straight-banded Treebrown

• Satyrus verma Kollar, [1844]; in Hügel, Kaschmir und das Reich der Siek 4: 447, pl. 16, f. 1-2

• Lethe (Lethe) verma; Lang, 2020, Atalanta 51 (3/4): 334 (list)
• Lethe verma; Moore, 1878, Proc. zool. Soc. Lond. 1878 (4): 824; Seitz, 1907, Gross-Schmett. Erde 1: 84, pl. 30 e; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 324; [BOW]: pl. 168, f. 5; [BOR], 420; [MRS], 332; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]

600x400 (~63Kb) upperside India: Arunachal Pradesh, Taley Valley WLS, 10.11.2005, Photo © Kedar Tokekar

600x400 (~47Kb) underside India: Arunachal Pradesh, Taley Valley WLS, 10.11.2005, Photo © Kedar Tokekar

• L. v. verma NW.India

  • Lethe verma verma; [BOR], 420; [iNaturalist]
• L. v. stenopa Fruhstorfer, 1908 Burma, Thailand - Indo China, S.China, Hainan

  • Lethe (Tausima) verma stenopa Fruhstorfer, 1908; Ent. Zs. 22 (31): 127; TL: N.Vietnam
  • ?Lethe (Tausima) verma stenopa f. laticincta Fruhstorfer, 1908; Ent. Zs. 22 (31): 127

  • Lethe verma stenopa; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 324; [BOR], 420; [MRS], 332; [iNaturalist] ; [Yutaka Inayoshi]
  • ?Lethe verma stenopa f. laticincta; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 324
• L. v. cintamani Fruhstorfer, 1909 Taiwan

  • Lethe verma cintamani Fruhstorfer, 1909; Ent. Zs. 23 (8): 40; TL: Formosa

  • Lethe verma cintamani; [BOR], 420; [MRS], 332; [iNaturalist]
• L. v. sintica Fruhstorfer, 1911 Sikkim, Assam, Yunnan

  • Lethe verma sintica Fruhstorfer, 1911; Gross-Schmett. Erde 9: 324

  • Lethe verma sintica; [BOR], 420; Huang, 2001; Huang, 2003, Neue Ent. Nachr. 55: 94 (note); [iNaturalist]
• L. v. satarnus Fruhstorfer, 1911 W.China

  • Lethe verma satarnus Fruhstorfer, 1911; Gross-Schmett. Erde 9: 324

  • Lethe verma satarnus; [BOR], 420; [MRS], 332
• L. v. robinsoni Pendlebury, 1933 Peninsular Malaya (Cameron Highlands)

  • Lethe robinsoni Pendlebury, 1933; J. Fed. Malay St. Mus. 17 (2): 386

  • Lethe verma robinsoni; [BOR], 420; [BMP]: 125, pl. 15, f. 13; [LepIndex]; [CoL]

Lethe confusa Aurivillius, 1897 Murree - Assam, Burma, Malaysia. See [maps] Banded Treebrown

• Lethe confusa Aurivillius, 1897; Ent. Tidskr. 18 (3/4): 142; TL: [India]
  • = Lethe rohria; Seitz, 1907, Gross-Schmett. Erde 1: 84

• Lethe (Lethe) confusa; Lang, 2020, Atalanta 51 (3/4): 334 (list)
• Lethe confusa; [EBW]; [BOW]: pl. 166, f. 15-16; [BOR], 420; [MRS], 331; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]

512x341 (~52Kb) underside India: Guwahati, 17.11.2005, Photo © Kedar Tokekar

Larva on Microstegium ciliatum, Bambusa multiplex [MRS]

• L. c. confusa N.India, Himalayas, Assam, Burma, Thailand, Indo-China, Malaya, SE.Tibet, S.China, Hainan, Yunnan

  • Lethe confusa fuhaica Lee, 1962; Acta ent. sin. 11 (2): 139, 144; TL: Yunnan

  • Lethe confusa confusa; [BOR], 420; Huang, 2003, Neue Ent. Nachr. 55: 94 (note); [iNaturalist] ; [Yutaka Inayoshi]
  • Lethe confusa fuhaica; Huang & Wu, 2003, Neue Ent. Nachr. 55: 126, pl. 10, f. 14; [LepIndex]; [CoL]

  1165x1392 (~240Kb) underside Cambodia, Pursat Province, Phnom Tumpor Mt., the O'Gran brook valley in tall forest, 1190 m a.s.l., 17th March 2019, Photo © Oleg Kosterin

  1442x1218 (~230Kb) underside Cambodia, Pursat Province, Phnom Tumpor Mt., the O'Gran brook valley in tall forest, 1190 m a.s.l., 17th March 2019, Photo © Oleg Kosterin

  1248x1155 (~349Kb) underside Cambodia, Pursat Province, Phnom Tumpor Mt., the O'Gran brook valley in tall forest, 1190 m a.s.l., 17th March 2019, Photo © Oleg Kosterin

• L. c. apara Fruhstorfer, 1911 S.Burma, Thailand - Indo China, Hong Kong, Hainan

  • Lethe rohria apara Fruhstorfer, 1911; Gross-Schmett. Erde 9: 315; TL: Tenasserim

  • Lethe confusa apara; [BOR], 420; [MRS], 331
• L. c. gambara Fruhstorfer, 1911 Sikkim, Assam

  • Lethe rohria gambara Fruhstorfer, 1911; Gross-Schmett. Erde 9: 315; TL: Assam

  • Lethe confusa gambara; [BOR], 420
• L. c. enima Fruhstorfer, 1911 Sumatra, Peninsular Malaya

  • Lehte rohria enima Fruhstorfer, 1911; Gross-Schmett. Erde 9: 315, pl. 97 a; TL: Sumatra

  • Lethe confusa enima; [BMP]: 125, pl. 15, f. 19; [BOR], 420; [iNaturalist]
• L. c. godana Fruhstorfer, 1911 Java

  • Lethe rothria godana Fruhstorfer, 1911; Gross-Schmett. Erde 9: 315; TL: Java

  • Lethe confusa godana; [BOR], 420; [iNaturalist]
• L. c. kalisi Lamas, 2010 Borneo

  • Lethe confusa kalisi Lamas, 2010; SHILAP Revta. lepid. 38 (150): 199 (repl. Lethe rohria borneensis Kalis, 1933); TL: Borneo
  • Lethe rohria borneensis Kalis, 1933; Tijdschr. Ent. 76 (1-2): 69 (preocc. Lethe darena var. borneensis Staudinger, [1897]); TL: Borneo

  • Lethe rohria borneensis; [LepIndex]; [CoL]

Lethe hyrania (Kollar, [1844]) W.China, Chamba - Assam. See [maps] Common Forester

• Satyrus hyrania Kollar, [1844]; in Hügel, Kaschmir und das Reich der Siek 4: 449, pl. 17, f. 1-2
  • = Lethe insana; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 317
• Satyrus isana Kollar, [1844]; in Hügel, Kaschmir und das Reich der Siek 4: 448, pl. 16, f. 3-4
  • = Lethe hyrania hyrania; Lang & Lamas, 2016, Zootaxa 4072 (3): 396

• Lethe (Lethe) hyrania; Lang, 2020, Atalanta 51 (3/4): 334 (list)
• Lethe hyrania; [iNaturalist]
• Lethe insana; [LepIndex] ; [CoL]
• Lethe insana[sic, isana]; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 317; [BOR], 420; [MRS], 331; Yoshino, 2008, Futao (54): 9, pl. 2, f. 5-6
• Lethe isana isana; [iNaturalist]
• Lethe isana; [BOW]: pl. 202, f. 2; [iNaturalist]

Larva on (as isana) Arundinaria falcata [MRS]

• L. h. hyrania NW.India

  • Lethe hyrania hyrania; Lang & Lamas, 2016, Zootaxa 4072 (3): 396
  • Lethe insana insana[sic, isana]; [BOR], 420
• L. h. dinarbas (Hewitson, 1863) China (NW.Yunnan), SE.Tibet, N.Burma, Sikkim, Bhutan, Nepal, N.India

  • Debis dinarbas Hewitson, 1863; Ill. exot. Butts [4] (Debis II-III): [37], pl. [20], f. 15; TL: North India

  • Lethe hyrania dinarbas; Lang & Lamas, 2016, Zootaxa 4072 (3): 397
  • Lethe insana dinarbas; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 317
  • Lethe isana dinarbas; [iNaturalist]
• L. h. formosana Fruhstorfer, 1908 Taiwan

  • Lethe insana formosana Fruhstorfer, 1908; Ent. Zs. 22 (12): 48; TL: Formosa, Larachi
  • Lethe baucis var. arisana Matsumura, 1909; Ent. Zs. 23 (19): 91; TL: Formosa
    • = ; Lang & Lamas, 2016, Zootaxa 4072 (3): 397 (nom. nud.)
  • Lethe insana formosana ab. uraiensis Murayama & Shimonoya, 1963; 21
    • = ; Lang & Lamas, 2016, Zootaxa 4072 (3): 397 (nom. nud.)

  • Lethe hyrania formosana; Lang & Lamas, 2016, Zootaxa 4072 (3): 397
  • Lethe insana formosana; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 318; [BOR], 420; [MRS], 331; [iNaturalist]
• L. h. caerulescens Mell, 1923 China (S.Anhui, Jiangxi, Fujian, Guangdong, Hainan, Guangxi), N.Vietnam, Laos

  • Lethe insana[sic] caerulescens Mell, 1923; Dt. ent. Zs. 1923 (2): 155; TL: Kuangtung

  • Lethe hyrania caerulescens; Lang & Lamas, 2016, Zootaxa 4072 (3): 397; [Yutaka Inayoshi]

Lethe baucis Leech, 1891 China (W.Hubei, Sichuan, Chongqing). See [maps]

• Lethe baucis Leech, 1891; Entomologist 24 (Suppl.): 3; TL: Wa-Shan; Chia-Kou-Ho; Ta-Tsien-Lu
• Lethe procris Leech, 1891; Entomologist 24 (Suppl.): 2; TL: Wa-Shan; Chang-Yang
  • = ; Lang, 2016, Phegea 44 (2): 35

• Lethe (Lethe) baucis; Lang, 2020, Atalanta 51 (3/4): 334 (list)
• Lethe baucis; Seitz, 1907, Gross-Schmett. Erde 1: 84, pl. 30 c; [BOW]: pl. 201, f. 8-9; Lang, 2016, Phegea 44 (2): 35; [LepIndex] ; [CoL] ; [iNaturalist] ; [Yutaka Inayoshi]
• Lethe baucis ab. procris; Seitz, 1907, Gross-Schmett. Erde 1: 84
• Lethe insana baucis; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 318; [MRS], 331
• Lethe insana procris; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 318

• L. b. huanghaoi Lang, 2016 N.Yunnan

  • Lethe baucis huanghaoi Lang, 2016; Phegea 44 (2): 37; TL: China, Yunnan, Binchuan, Mt Juzishan, Muxaingping, 2300m

Lethe marginalis Motschulsky, 1860 E.China, Amur, Korea, Japan. See [maps]

• Lethe marginalis Motschulsky, 1860; Etud. Ent. 9: 29; TL: Hakodate, Hokkaido, Japan
• Satyrus (Pararge) davidianus Poujade, 1885; Bull. Soc. Ent. Fr. (6) 5: xciv; TL: Mou-Pin
  • = Lethe lanaris; Seitz, 1907, Gross-Schmett. Erde 1: 83

• Lethe (Lethe) marginalis; Lang, 2020, Atalanta 51 (3/4): 334 (list)
• Lethe marginalis; Seitz, 1907, Gross-Schmett. Erde 1: 86, pl. 31 d; Kudrna, 1974, Atalanta 5: 101; [BRU], 184; [BOW]: pl. 202, f. 4; [MRS], 338; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 44; [LepIndex] ; [CoL] ; [iNaturalist]

Larva on Miscanthus sinensis, Spodiopogon sibiricus, Scirpus wichurae, Carex [BRU]

• L. m. marginalis

  • Lethe marginalis marginalis; Huang, 2003, Neue Ent. Nachr. 55: 94 (note), f. 151
• L. m. maackii (Bremer, 1861) Amur, Ussuri

  • Lasiommata maackii Bremer, 1861; Bull. Acad. Imp. Sci. St. Petersb. 3: 468, Mélanges biol. St.-Pétersb. 3 (5-6):549
    • = Lethe marginalis; Seitz, 1907, Gross-Schmett. Erde 1: 86

  • Lethe marginalis maacki; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 44

  824x855 (~232Kb) upperside Russia, Khabarovskiy Kray, Bolshekhekhtsirskiy Natural Reserve, Chirki post. 15th July 2014, Photo © Oleg Kosterin

• L. m. obscurofasciata Huang, 2002 Yunnan

  • Lethe marginalis obscurofasciata Huang, 2002; Atalanta 33 (3/4): 369, pl. 22, f. 2, 6; TL: Gazu, Nujiang valley, Yunnan, 1800m

  • Lethe marginalis obscurofasciata; Huang, 2003, Neue Ent. Nachr. 55: 94 (note), f. 150

Lethe kansa (Moore, 1857) Kumaon - Assam, Burma. See [maps] Bamboo Forester

• Debis kansa Moore, 1857; in Horsfield & Moore, Cat. lep. Ins. Mus. East India Coy 1: 220; TL: Darjeeling

• Lethe (Lethe) kansa; Lang, 2020, Atalanta 51 (3/4): 334 (list)
• Lethe kansa; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 318; [BOW]: pl. 167, f. 7; [BOR], 419; [MRS], 324; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [Yutaka Inayoshi]

• L. k. kansa Sikkim

  • Lethe kansa kansa; [BOR], 419
• L. k. zeugitana Fruhstorfer, 1911 Assam, Manipur

  • Lethe kansa zeugitana Fruhstorfer, 1911; Gross-Schmett. Erde 9: 318, pl. 98 a (as kansa); TL: Assam

  • Lethe kansa zeugitana; [BOR], 419
• L. k. vaga Fruhstorfer, 1911 S.Burma, S.Thailand, S.Yunnan

  • Lethe kansa vaga Fruhstorfer, 1911; Gross-Schmett. Erde 9: 318-319; TL: Tenasserim

  • Lethe kansa vaga; [BOR], 419; [MRS], 324

Lethe sinorix (Hewitson, 1863) China, Sikkim - Assam, S.Burma, Malaya. See [maps] Red Tailed Forester

• Debis sinorix Hewitson, 1863; Ill. exot. Butts [4] (Debis II-III): [38], pl. [20], f. 19-20; TL: East India
• Lethe sinorix kuangtungensis Mell, 1923; Dt. ent. Zs. 1923 (2): 154; TL: Kuangtung
• lotaoshanensis Tinkham, 1936; Lingman J. Sci. 15: 561

• Lethe (Lethe) sinorix; Lang, 2020, Atalanta 51 (3/4): 334 (list)
• Lethe sinorix lotaoshanensis; [LepIndex]; [CoL]
• Lethe sinorix; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 319, pl. 98 b; [BOW]: pl. 168, f. 2; [BOR], 418; [MRS], 324; [LepIndex] ; [CoL] ; [iNaturalist]

• L. s. sinorix Sikkim, Bhutan, Assam, Burma, ?Thailand

  • Lethe sinorix sinorix; [BOR], 418; [Yutaka Inayoshi]
• L. s. vanda Corbet, 1941 Peninsular Malaya (Cameron Highlands)

  • Lethe vanda Corbet, 1941; J. fed. Malay St. Mus. 18 (5): 807

  • Lehte sinorix vanda; [BOR], 419
  • Lethe sinorix vanda; [BMP]: 125, pl. 15, f. 10; [LepIndex]; [CoL]
• L. s. obscura Mell, 1942

  • Lethe sinorix obscura Mell, 1942; Arch. Naturgesch. (N.F.) 11: 274; TL: N.Fukien, Kuatun
  • Lethe sinorix xironis Lamas, 2010; SHILAP Revta. lepid. 38 (150): 200 (repl. Lethe sinorix obscura Mell, 1942)

  • Lethe sinorix obscura; [MRS], 324; [LepIndex]; [CoL]

Lethe minerva (Fabricius, 1775) Malaya, Sumatra, Borneo, Java. See [maps]

• Papilio minerva Fabricius, 1775; Syst. Ent.: 493, no. 216
• Papilio arcadia Cramer, [1777]; Uitl. Kapellen 2 (9-16): pl. 116, f. E, F
  • = ; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 318
• Satyrus caumas Godart, [1824]; Encyclopédie Méthodique 9 (2): 479
  • = ; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 318

• Lethe (Lethe) minerva; Lang, 2020, Atalanta 51 (3/4): 334 (list)
• Lethe arcadia; Moore, 1878, Proc. zool. Soc. Lond. 1878 (4): 824
• Lethe minerva; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 318, pl. 98 a; [BOW]: pl. 167, f. 10; [BOR], 421; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]
• Nemetis minerva; [EBW]

• L. m. minerva Java, Sumatra, Peninsular Malaya

  • Lethe minerva minerva; [BMP]: 125, pl. 15, f. 12; [BOR], 421
• L. m. tritogeneia Fruhstorfer, 1911 Burma, Thailand

  • Lethe minerva tritogeneia Fruhstorfer, 1911; Gross-Schmett. Erde 9: 318; TL: Tenasserim, Myanmar

  • Lethe minerva tritogeneia; [BOR], 421; [Yutaka Inayoshi]

  1082x1250 (~402Kb) ♂ underside Cambodia, Ratanakiri Province, Lake Yak Lom, 30th May 2013, Photo © Oleg Kosterin

Lethe chandica (Moore, [1858]) W.China, Assam - Burma, Malaya. See [maps] Angled Red Forester

• Debis chandica Moore, [1858]; in Horsfield & Moore, Cat. lep. Ins. Mus. East India Coy 1: 219; TL: Darjeeling, India
• Debis isabella Butler, 1866; Proc. zool. Soc. Lond. 1866 (1): 41, pl. 3, f. 4; TL: Philippines
• Lethe chandica obscura Mell, 1942; Arch. Naturgesch. (N.F.) 11: 276 (preocc. Lethe callipteris obscura Nakahara, 1926)
• Lethe chandica f. rahula Fruhstorfer, 1911; Gross-Schmett. Erde 9: 320; TL: Hainan

• Lethe (Lethe) chandica; Lang, 2020, Atalanta 51 (3/4): 334 (list)
• Lethe chandica obscura; [LepIndex]; [CoL]
• Lethe chandica; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 320; [BOW]: pl. 166, f. 11; [BOR], 422; [MRS], 329; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]

512x341 (~36Kb) underside India: Itanagar, 13.11.2005, Photo © Kedar Tokekar

Larva on Phyllostachys pubescens [MRS]

• L. c. chandica NW.India - Peninsular Malaya?, Sumatra?, Yunnan

  • Lethe chandica chandica; [BOR], 422; Huang, 2003, Neue Ent. Nachr. 55: 94 (note); [iNaturalist] ; [Yutaka Inayoshi]
• L. c. negrito (C. & R. Felder, 1863) Philippines (Luzon)

  • Debis negrito C. & R. Felder, 1863; Wien. ent. Monats. 7 (4): 124; TL: Luzon

  • Lethe chandica negrito; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 320; [BOR], 422
• L. c. coelestis Leech, [1892] W.China

  • Lethe chandica var. coelestis Leech, [1892]; Butts China Japan Corea (1): 20, pl. 3, f. 7-8

  • Lethe chandica coelestis; Seitz, 1907, Gross-Schmett. Erde 1: 84, pl. 31 a; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 320; [BOR], 422; [MRS], 329
• L. c. ratnacri Fruhstorfer, 1908 Taiwan

  • Lethe chandica ratnacri Fruhstorfer, 1908; Ent. Wochenbl. 25 (9): 38; TL: Formosa, Kagi

  • Lethe chandica ratnacri; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 320, pl. 98 b; [BOR], 422; [MRS], 329; [iNaturalist]
• L. c. suvarna Fruhstorfer, 1908 Laos, N.Vietnam, Hainan

  • Lethe chandica suvarna Fruhstorfer, 1908; Ent. Wochenbl. 25 (9): 38
  • Letha chandica suvarna f. suvarna Fruhstorfer, 1908; Ent. Wochenbl. 25 (9): 38
  • Letha chandica suvarna f. rahula Fruhstorfer, 1908; Ent. Wochenbl. 25 (9): 38
  • Lethe chandica suvarna Fruhstorfer, 1911; Gross-Schmett. Erde 9: 320; TL: Chiem Hoa, N.Vietnam

  • Lethe chandica suvarna; [MRS], 329; [iNaturalist] ; [Yutaka Inayoshi]
• L. c. namura Fruhstorfer, 1911 Peninsular Malaya, Sumatra?

  • Lethe chandica namura Fruhstorfer, 1911; Gross-Schmett. Erde 9: 321

  • Lethe chandica namura; [BMP]: 125, pl. 15, f. 15, 17; [Yutaka Inayoshi]
• L. c. marga Fruhstorfer, 1911 Java

  • Lethe chandica marga Fruhstorfer, 1911; Gross-Schmett. Erde 9: 321; TL: W.Java

  • Lethe chandica marga; [BOR], 422
• L. c. flanona Fruhstorfer, 1911 Assam

  • Lethe chandica flanona Fruhstorfer, 1911; Gross-Schmett. Erde 9: 320; TL: Assam

  • Lethe chandica flanona; [MRS], 329; [iNaturalist]
• L. c. sisapon Fruhstorfer, 1911 Philippines (Mindoro)

  • Lethe chandica sisapon Fruhstorfer, 1911; Gross-Schmett. Erde 9: 321; TL: Mindoro

  • Lethe chandica sisapon; [BOR], 422
• L. c. byzaccus Fruhstorfer, 1911 Philippines (Mindanao)

  • Lethe chandica byzaccus Fruhstorfer, 1911; Gross-Schmett. Erde 9: 321; TL: Mindanao

  • Lethe chandica byzaccus; [BOR], 422
• L. c. jomaria Fruhstorfer, 1911 Sulu Is.

  • Lethe chandica jomaria Fruhstorfer, 1911; Gross-Schmett. Erde 9: 321; TL: Jolo

  • Lethe chandica jomaria; [BOR], 422
• L. c. ratnapandi Fruhstorfer, 1911 Palawan

  • Lethe chandica ratnapandi Fruhstorfer, 1911; Gross-Schmett. Erde 9: 321; TL: Palawan

  • Lethe chandica ratnapandi; [BOR], 422
  • ?Lethe negrito var.; Staudinger, 1889, Dt. ent. Z. Iris 2 (1): 34

Lethe mekara (Moore, [1858]) China, Assam - Burma, Malaya. See [maps] Common Red Forester

• Debis mekara Moore, [1858]; in Horsfield & Moore, Cat. lep. Ins. Mus. East India Coy 1: 219; TL: Darjeeling
• Lethe mekara f. vajra Fruhstorfer, 1911; Gross-Schmett. Erde 9: 319
• manis Corbet, 1942; Ann. Mag. nat. Hist. (11) 9 (56): 609; TL: Mentawei I.

• Lethe (Lethe) mekara; Lang, 2020, Atalanta 51 (3/4): 334 (list)
• Lethe mekara manis; [LepIndex]; [CoL]
• Lethe mekara; Moore, 1878, Proc. zool. Soc. Lond. 1878 (4): 824; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 319; [BOW]: pl. 167, f. 10; [BOR], 422; [MRS], 330; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM]

Larva on Phyllostachys pubescens [MRS]

• L. m. mekara N.India, Sikkim

  • Lethe mekara mekara; [BOR], 422
• L. m. zuchara Fruhstorfer, 1911 Assam - Tenasserim, Indo China

  • Lethe mekara zuchara Fruhstorfer, 1911; Gross-Schmett. Erde 9: 319

  • Lethe mekara zuchara; [BOR], 422; [iNaturalist]
• L. m. crijnana Fruhstorfer, 1911 Thailand, Laos, N.Vietnam, S.Yunnan

  • Lethe mekara crijnana Fruhstorfer, 1911; Gross-Schmett. Erde 9: 319, pl. 98 c; TL: N.Vietnam

  • Lethe mekara crijnana; [MRS], 330; [Yutaka Inayoshi]

  1112x1250 (~327Kb) ♂ underside Cambodia, Koh Kong Province, Stung Kep ("Left Tatai") River valley at Rainbow Lodge, bamboo forest, 24th August 2011, Photo © Oleg Kosterin

• L. m. gopaka Fruhstorfer, 1911 Peninsular Malaya

  • Lethe mekara gopaka Fruhstorfer, 1911; Gross-Schmett. Erde 9: 320; TL: Malaysia

  • Lethe mekara gopaka; [BOR], 422; [BMP]: 125, pl. 15, f. 14, 16, 18; [Yutaka Inayoshi]
• L. m. debata Fruhstorfer, 1911 Sumatra

  • Lethe mekara debata Fruhstorfer, 1911; Gross-Schmett. Erde 9: 320, pl. 98 d; TL: Celebes

  • Lethe mekara debata; [BOR], 422
• L. m. sumati Fruhstorfer, 1911 Borneo

  • Lethe mekara sumati Fruhstorfer, 1911; Gross-Schmett. Erde 9: 319, pl. 98 c d; TL: N.Borneo

  • Lethe mekara sumati; [BOR], 422

Lethe vindhya (C. & R. Felder, 1859) Sikkim - Burma, Assam - Tenasserim. See [maps] Black Forester

• Debis vindhya C. & R. Felder, 1859; Wien. ent. Monats. 3 (12): 402; TL: Assam
• Lethe alberta Butler, 1871; Ann. Mag. nat. Hist. (4) 8 (46): 283; TL: "Benares"
• Lethe dolopes Hewitson, 1872; Ent. mon. Mag. 9: 85; TL: Darjeling

• Lethe (Lethe) vindhya; Lang, 2020, Atalanta 51 (3/4): 334 (list)
• Lethe vindhya dolopes; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 319
• Lethe vindhya vindhya; [iNaturalist]
• Lethe vindhya; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 319, pl. 98 b; [BOR], 424; [MRS], 339; [LepIndex] ; [CoL] ; [iNaturalist]

Larva on Bambusa spp. [MRS]

• L. v. ladesta Fruhstorfer, 1911

  • Lethe vindhya ladesta Fruhstorfer, 1911; Gross-Schmett. Erde 9: 319; TL: Tenasserim

  • Lethe vindhya ladesta; [MRS], 339
• L. v. luaba Corbet, 1941

  • Lethe luaba Corbet, 1941; J. fed. Malay St. Mus. 18 (5): 807

  • Lethe vindhya luaba; [BMP]: 125, pl. 15, f. 11; [LepIndex]; [CoL]

Lethe serbonis (Hewitson, 1876) Bhutan, Sikkim, Assam - W.China. See [maps] Brown Forester

• Debis serbonis Hewitson, 1876; Ent. mon. Mag. 13 (7): 151; TL: Darjeeling
• Debis davidi Oberthür, 1881; Étud. d'Ent. 6: 15, pl. 7, f. 5; TL: Mou-Pin
• Lethe davidi var. flavofasciata Leech, [1892]; Butts China Japan Corea (1): 30
• Lethe serbonis naganum Tytler, 1914; J. Bombay nat. Hist. Soc. 23 (2): 219; TL: Kabru Peak, Manipur; Naga Hills
• teesta Talbot, 1947; Fauna Br. Ind., Butt. (2nd ed.) 2: 220; TL: Sikkim
• bhutya Talbot, 1947; Fauna Br. Ind., Butt. (2nd ed.) 2: 221; TL: Bhutan
• Lethe serbonis davidi f. angulata Seitz, 1907; Gross-Schmett. Erde 1: 83

• Lethe (Lethe) serbonis; Lang, 2020, Atalanta 51 (3/4): 334 (list)
• Lethe serbonis bhutya; [LepIndex]; [CoL]
• Lethe serbonis davidi; Seitz, 1907, Gross-Schmett. Erde 1: 83, pl. 30 b; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 319
• Lethe serbonis davidi ab. flavofasciata; Seitz, 1907, Gross-Schmett. Erde 1: 83
• Lethe serbonis davidi f. flavofasciata; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 319
• Lethe serbonis teesta; [LepIndex]; [CoL]
• Lethe serbonis; Fruhstorfer, 1911, Gross-Schmett. Erde 9: 319, pl. 98 b; [BOW]: pl. 167, f. 9; [BOR], 424; [MRS], 330; [LepIndex] ; [CoL] ; [iNaturalist]

• ?L. s. pallida Tytler, 1939 Burma

  • Lethe serbonis pallida Tytler, 1939; J. Bombay nat. Hist. Soc. 41 (2): 245; TL: NE.Burma

  • Lethe serbonis pallida; Huang, 2001; [LepIndex]; [CoL]
• L. s. gesala Lang, 2014

  • Lethe serbonis gesala Lang, 2014; Neue Ent. Nachr. 70: (299-308)